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def load_multiple(filters="*.*", text='Select some files, FACEFACE!', default_directory='default_directory'): """ Pops up a dialog for opening more than one file. Returns a list of string paths or None. """ # make sure the filters contains "*.*" as an option! if not '*' in filters.split(';'): filters = filters + ";;All files (*)" # if this type of pref doesn't exist, we need to make a new one if default_directory in _settings.keys(): default = _settings[default_directory] else: default = "" # pop up the dialog results = _qtw.QFileDialog.getOpenFileNames(None,text,default,filters) # If Qt5, take the zeroth element if _s._qt.VERSION_INFO[0:5] == "PyQt5": results = results[0] # Make sure it's a string result = [] for r in results: result.append(str(r)) if len(result)==0: return else: _settings[default_directory] = _os.path.split(result[0])[0] return result
Pops up a dialog for opening more than one file. Returns a list of string paths or None.
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def Dump(self): """ Dumps the current prefs to the preferences.txt file """ prefs_file = open(self.prefs_path, 'w') for n in range(0,len(self.prefs)): if len(list(self.prefs.items())[n]) > 1: prefs_file.write(str(list(self.prefs.items())[n][0]) + ' = ' + str(list(self.prefs.items())[n][1]) + '\n') prefs_file.close()
Dumps the current prefs to the preferences.txt file
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def link(self): """Resolve and link all types in the scope.""" type_specs = {} types = [] for name, type_spec in self.scope.type_specs.items(): type_spec = type_spec.link(self.scope) type_specs[name] = type_spec if type_spec.surface is not None: self.scope.add_surface(name, type_spec.surface) types.append(type_spec.surface) self.scope.type_specs = type_specs self.scope.add_surface('__types__', tuple(types))
Resolve and link all types in the scope.
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def install(path, name=None): """Compiles a Thrift file and installs it as a submodule of the caller. Given a tree organized like so:: foo/ __init__.py bar.py my_service.thrift You would do, .. code-block:: python my_service = thriftrw.install('my_service.thrift') To install ``my_service`` as a submodule of the module from which you made the call. If the call was made in ``foo/bar.py``, the compiled Thrift file will be installed as ``foo.bar.my_service``. If the call was made in ``foo/__init__.py``, the compiled Thrift file will be installed as ``foo.my_service``. This allows other modules to import ``from`` the compiled module like so, .. code-block:: python from foo.my_service import MyService .. versionadded:: 0.2 :param path: Path of the Thrift file. This may be an absolute path, or a path relative to the Python module making the call. :param str name: Name of the submodule. Defaults to the basename of the Thrift file. :returns: The compiled module """ if name is None: name = os.path.splitext(os.path.basename(path))[0] callermod = inspect.getmodule(inspect.stack()[1][0]) name = '%s.%s' % (callermod.__name__, name) if name in sys.modules: return sys.modules[name] if not os.path.isabs(path): callerfile = callermod.__file__ path = os.path.normpath( os.path.join(os.path.dirname(callerfile), path) ) sys.modules[name] = mod = load(path, name=name) return mod
Compiles a Thrift file and installs it as a submodule of the caller. Given a tree organized like so:: foo/ __init__.py bar.py my_service.thrift You would do, .. code-block:: python my_service = thriftrw.install('my_service.thrift') To install ``my_service`` as a submodule of the module from which you made the call. If the call was made in ``foo/bar.py``, the compiled Thrift file will be installed as ``foo.bar.my_service``. If the call was made in ``foo/__init__.py``, the compiled Thrift file will be installed as ``foo.my_service``. This allows other modules to import ``from`` the compiled module like so, .. code-block:: python from foo.my_service import MyService .. versionadded:: 0.2 :param path: Path of the Thrift file. This may be an absolute path, or a path relative to the Python module making the call. :param str name: Name of the submodule. Defaults to the basename of the Thrift file. :returns: The compiled module
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def loads(self, name, document): """Parse and compile the given Thrift document. :param str name: Name of the Thrift document. :param str document: The Thrift IDL as a string. """ return self.compiler.compile(name, document).link().surface
Parse and compile the given Thrift document. :param str name: Name of the Thrift document. :param str document: The Thrift IDL as a string.
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def load(self, path, name=None): """Load and compile the given Thrift file. :param str path: Path to the ``.thrift`` file. :param str name: Name of the generated module. Defaults to the base name of the file. :returns: The compiled module. """ if name is None: name = os.path.splitext(os.path.basename(path))[0] # TODO do we care if the file extension is .thrift? with open(path, 'r') as f: document = f.read() return self.compiler.compile(name, document, path).link().surface
Load and compile the given Thrift file. :param str path: Path to the ``.thrift`` file. :param str name: Name of the generated module. Defaults to the base name of the file. :returns: The compiled module.
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def get_url_implicit_flow_user(client_id, scope, redirect_uri='https://oauth.vk.com/blank.html', display='page', response_type='token', version=None, state=None, revoke=1): """ https://vk.com/dev/implicit_flow_user :return: url """ url = "https://oauth.vk.com/authorize" params = { "client_id": client_id, "scope": scope, "redirect_uri": redirect_uri, "display": display, "response_type": response_type, "version": version, "state": state, "revoke": revoke } params = {key: value for key, value in params.items() if value is not None} return u"{url}?{params}".format(url=url, params=urlencode(params))
https://vk.com/dev/implicit_flow_user :return: url
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def get_url_authcode_flow_user(client_id, redirect_uri, display="page", scope=None, state=None): """Authorization Code Flow for User Access Token Use Authorization Code Flow to run VK API methods from the server side of an application. Access token received this way is not bound to an ip address but set of permissions that can be granted is limited for security reasons. Args: client_id (int): Application id. redirect_uri (str): Address to redirect user after authorization. display (str): Sets authorization page appearance. Sets: {`page`, `popup`, `mobile`} Defaults to `page` scope (:obj:`str`, optional): Permissions bit mask, to check on authorization and request if necessary. More scope: https://vk.com/dev/permissions state (:obj:`str`, optional): An arbitrary string that will be returned together with authorization result. Returns: str: Url Examples: >>> vk.get_url_authcode_flow_user(1, 'http://example.com/', scope="wall,email") 'https://oauth.vk.com/authorize?client_id=1&display=page&redirect_uri=http://example.com/&scope=wall,email&response_type=code .. _Docs: https://vk.com/dev/authcode_flow_user """ url = "https://oauth.vk.com/authorize" params = { "client_id": client_id, "redirect_uri": redirect_uri, "display": display, "response_type": "code" } if scope: params['scope'] = scope if state: params['state'] = state return u"{url}?{params}".format(url=url, params=urlencode(params))
Authorization Code Flow for User Access Token Use Authorization Code Flow to run VK API methods from the server side of an application. Access token received this way is not bound to an ip address but set of permissions that can be granted is limited for security reasons. Args: client_id (int): Application id. redirect_uri (str): Address to redirect user after authorization. display (str): Sets authorization page appearance. Sets: {`page`, `popup`, `mobile`} Defaults to `page` scope (:obj:`str`, optional): Permissions bit mask, to check on authorization and request if necessary. More scope: https://vk.com/dev/permissions state (:obj:`str`, optional): An arbitrary string that will be returned together with authorization result. Returns: str: Url Examples: >>> vk.get_url_authcode_flow_user(1, 'http://example.com/', scope="wall,email") 'https://oauth.vk.com/authorize?client_id=1&display=page&redirect_uri=http://example.com/&scope=wall,email&response_type=code .. _Docs: https://vk.com/dev/authcode_flow_user
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def get_fake_data(*a): """ Called whenever someone presses the "fire" button. """ # add columns of data to the databox d['x'] = _n.linspace(0,10,100) d['y'] = _n.cos(d['x']) + 0.1*_n.random.rand(100) # update the curve c.setData(d['x'], d['y'])
Called whenever someone presses the "fire" button.
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def setOpts(self, **opts): """ Changes the behavior of the SpinBox. Accepts most of the arguments allowed in :func:`__init__ <pyqtgraph.SpinBox.__init__>`. """ #print opts for k in opts: if k == 'bounds': #print opts[k] self.setMinimum(opts[k][0], update=False) self.setMaximum(opts[k][1], update=False) #for i in [0,1]: #if opts[k][i] is None: #self.opts[k][i] = None #else: #self.opts[k][i] = D(unicode(opts[k][i])) elif k in ['step', 'minStep']: self.opts[k] = D(asUnicode(opts[k])) elif k == 'value': pass ## don't set value until bounds have been set else: self.opts[k] = opts[k] if 'value' in opts: self.setValue(opts['value']) ## If bounds have changed, update value to match if 'bounds' in opts and 'value' not in opts: self.setValue() ## sanity checks: if self.opts['int']: if 'step' in opts: step = opts['step'] ## not necessary.. #if int(step) != step: #raise Exception('Integer SpinBox must have integer step size.') else: self.opts['step'] = int(self.opts['step']) if 'minStep' in opts: step = opts['minStep'] if int(step) != step: raise Exception('Integer SpinBox must have integer minStep size.') else: ms = int(self.opts.get('minStep', 1)) if ms < 1: ms = 1 self.opts['minStep'] = ms if 'delay' in opts: self.proxy.setDelay(opts['delay']) self.updateText()
Changes the behavior of the SpinBox. Accepts most of the arguments allowed in :func:`__init__ <pyqtgraph.SpinBox.__init__>`.
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def setMaximum(self, m, update=True): """Set the maximum allowed value (or None for no limit)""" if m is not None: m = D(asUnicode(m)) self.opts['bounds'][1] = m if update: self.setValue()
Set the maximum allowed value (or None for no limit)
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def selectNumber(self): """ Select the numerical portion of the text to allow quick editing by the user. """ le = self.lineEdit() text = asUnicode(le.text()) if self.opts['suffix'] == '': le.setSelection(0, len(text)) else: try: index = text.index(' ') except ValueError: return le.setSelection(0, index)
Select the numerical portion of the text to allow quick editing by the user.
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def value(self): """ Return the value of this SpinBox. """ if self.opts['int']: return int(self.val) else: return float(self.val)
Return the value of this SpinBox.
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def setValue(self, value=None, update=True, delaySignal=False): """ Set the value of this spin. If the value is out of bounds, it will be clipped to the nearest boundary. If the spin is integer type, the value will be coerced to int. Returns the actual value set. If value is None, then the current value is used (this is for resetting the value after bounds, etc. have changed) """ if value is None: value = self.value() bounds = self.opts['bounds'] if bounds[0] is not None and value < bounds[0]: value = bounds[0] if bounds[1] is not None and value > bounds[1]: value = bounds[1] if self.opts['int']: value = int(value) value = D(asUnicode(value)) if value == self.val: return prev = self.val self.val = value if update: self.updateText(prev=prev) self.sigValueChanging.emit(self, float(self.val)) ## change will be emitted in 300ms if there are no subsequent changes. if not delaySignal: self.emitChanged() return value
Set the value of this spin. If the value is out of bounds, it will be clipped to the nearest boundary. If the spin is integer type, the value will be coerced to int. Returns the actual value set. If value is None, then the current value is used (this is for resetting the value after bounds, etc. have changed)
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def interpret(self): """Return value of text. Return False if text is invalid, raise exception if text is intermediate""" strn = self.lineEdit().text() suf = self.opts['suffix'] if len(suf) > 0: if strn[-len(suf):] != suf: return False #raise Exception("Units are invalid.") strn = strn[:-len(suf)] try: val = fn.siEval(strn) except: #sys.excepthook(*sys.exc_info()) #print "invalid" return False #print val return val
Return value of text. Return False if text is invalid, raise exception if text is intermediate
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def editingFinishedEvent(self): """Edit has finished; set value.""" #print "Edit finished." if asUnicode(self.lineEdit().text()) == self.lastText: #print "no text change." return try: val = self.interpret() except: return if val is False: #print "value invalid:", str(self.lineEdit().text()) return if val == self.val: #print "no value change:", val, self.val return self.setValue(val, delaySignal=False)
Edit has finished; set value.
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def make_factory(self, cls, count): """ Get the generators from the Scaffolding class within the model. """ field_names = cls._meta.get_all_field_names() fields = {} text = [] finalizer = None scaffold = scaffolding.scaffold_for_model(cls) for field_name in field_names: generator = getattr(scaffold, field_name, None) if generator: if hasattr(generator, 'set_up'): generator.set_up(cls, count) fields[field_name] = generator text.append(u'%s: %s; ' % (field_name, fields[field_name])) try: self.stdout.write(u'Generator for %s: %s\n' % (cls, u''.join(text))) except models.ObjectDoesNotExist: self.stdout.write(u'Generator for %s\n' % u''.join(text)) if hasattr(scaffold, 'finalize') and hasattr(scaffold.finalize, '__call__'): finalizer = scaffold.finalize return fields, finalizer
Get the generators from the Scaffolding class within the model.
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def compute_indel_length(fs_df): """Computes the indel length accounting for wether it is an insertion or deletion. Parameters ---------- fs_df : pd.DataFrame mutation input as dataframe only containing indel mutations Returns ------- indel_len : pd.Series length of indels """ indel_len = pd.Series(index=fs_df.index) indel_len[fs_df['Reference_Allele']=='-'] = fs_df['Tumor_Allele'][fs_df['Reference_Allele']=='-'].str.len() indel_len[fs_df['Tumor_Allele']=='-'] = fs_df['Reference_Allele'][fs_df['Tumor_Allele']=='-'].str.len() indel_len = indel_len.fillna(0).astype(int) return indel_len
Computes the indel length accounting for wether it is an insertion or deletion. Parameters ---------- fs_df : pd.DataFrame mutation input as dataframe only containing indel mutations Returns ------- indel_len : pd.Series length of indels
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def keep_indels(mut_df, indel_len_col=True, indel_type_col=True): """Filters out all mutations that are not indels. Requires that one of the alleles have '-' indicating either an insertion or deletion depending if found in reference allele or somatic allele columns, respectively. Parameters ---------- mut_df : pd.DataFrame mutation input file as a dataframe in standard format indel_len_col : bool whether or not to add a column indicating the length of the indel Returns ------- mut_df : pd.DataFrame mutations with only frameshift mutations kept """ # keep only frameshifts mut_df = mut_df[is_indel_annotation(mut_df)] if indel_len_col: # calculate length mut_df.loc[:, 'indel len'] = compute_indel_length(mut_df) if indel_type_col: is_ins = mut_df['Reference_Allele']=='-' is_del = mut_df['Tumor_Allele']=='-' mut_df['indel type'] = '' mut_df.loc[is_ins, 'indel type'] = 'INS' mut_df.loc[is_del, 'indel type'] = 'DEL' return mut_df
Filters out all mutations that are not indels. Requires that one of the alleles have '-' indicating either an insertion or deletion depending if found in reference allele or somatic allele columns, respectively. Parameters ---------- mut_df : pd.DataFrame mutation input file as a dataframe in standard format indel_len_col : bool whether or not to add a column indicating the length of the indel Returns ------- mut_df : pd.DataFrame mutations with only frameshift mutations kept
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def keep_frameshifts(mut_df, indel_len_col=True): """Filters out all mutations that are not frameshift indels. Requires that one of the alleles have '-' indicating either an insertion or deletion depending if found in reference allele or somatic allele columns, respectively. Parameters ---------- mut_df : pd.DataFrame mutation input file as a dataframe in standard format indel_len_col : bool whether or not to add a column indicating the length of the frameshift Returns ------- mut_df : pd.DataFrame mutations with only frameshift mutations kept """ # keep only frameshifts mut_df = mut_df[is_frameshift_annotation(mut_df)] if indel_len_col: # calculate length mut_df.loc[:, 'indel len'] = compute_indel_length(mut_df) return mut_df
Filters out all mutations that are not frameshift indels. Requires that one of the alleles have '-' indicating either an insertion or deletion depending if found in reference allele or somatic allele columns, respectively. Parameters ---------- mut_df : pd.DataFrame mutation input file as a dataframe in standard format indel_len_col : bool whether or not to add a column indicating the length of the frameshift Returns ------- mut_df : pd.DataFrame mutations with only frameshift mutations kept
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def is_frameshift_len(mut_df): """Simply returns a series indicating whether each corresponding mutation is a frameshift. This is based on the length of the indel. Thus may be fooled by frameshifts at exon-intron boundaries or other odd cases. Parameters ---------- mut_df : pd.DataFrame mutation input file as a dataframe in standard format Returns ------- is_fs : pd.Series pandas series indicating if mutaitons are frameshifts """ # calculate length, 0-based coordinates #indel_len = mut_df['End_Position'] - mut_df['Start_Position'] if 'indel len' in mut_df.columns: indel_len = mut_df['indel len'] else: indel_len = compute_indel_length(mut_df) # only non multiples of 3 are frameshifts is_fs = (indel_len%3)>0 # make sure no single base substitutions are counted is_indel = (mut_df['Reference_Allele']=='-') | (mut_df['Tumor_Allele']=='-') is_fs[~is_indel] = False return is_fs
Simply returns a series indicating whether each corresponding mutation is a frameshift. This is based on the length of the indel. Thus may be fooled by frameshifts at exon-intron boundaries or other odd cases. Parameters ---------- mut_df : pd.DataFrame mutation input file as a dataframe in standard format Returns ------- is_fs : pd.Series pandas series indicating if mutaitons are frameshifts
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def get_frameshift_lengths(num_bins): """Simple function that returns the lengths for each frameshift category if `num_bins` number of frameshift categories are requested. """ fs_len = [] i = 1 tmp_bins = 0 while(tmp_bins<num_bins): if i%3: fs_len.append(i) tmp_bins += 1 i += 1 return fs_len
Simple function that returns the lengths for each frameshift category if `num_bins` number of frameshift categories are requested.
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def _init_context(self, gene_seq): """Initializes attributes defining mutation contexts and their position. The self.context2pos and self.pos2context dictionaries map from sequence context to sequence position and sequence position to sequence context, respectively. These attributes allow for randomly sampling of mutation positions while respecting sequence context in the randomization-based test. Parameters ---------- gene_seq : GeneSequence GeneSequence object from the gene_sequence module """ self.context2pos, self.pos2context = {}, {} gene_len = len(gene_seq.exon_seq) # get length of CDS five_ss_len = 2*len(gene_seq.five_prime_seq) # total length of 5' splice sites three_ss_len = 2*len(gene_seq.three_prime_seq) # total length of 3' splice sites if gene_seq.nuc_context in [1, 2]: # case where context matters index_context = int(gene_seq.nuc_context) - 1 # subtract 1 since python is zero-based index for i in range(index_context, gene_len): nucs = gene_seq.exon_seq[i-index_context:i+1] self.context2pos.setdefault(nucs, []) self.context2pos[nucs].append(i) self.pos2context[i] = nucs # sequence context for five prime splice site for i, five_ss in enumerate(gene_seq.five_prime_seq): first_nucs = five_ss[1-index_context:1+1] second_nucs = five_ss[2-index_context:2+1] first_pos = 2*i + gene_len second_pos = 2*i + gene_len + 1 self.context2pos.setdefault(first_nucs, []) self.context2pos[first_nucs].append(first_pos) self.context2pos.setdefault(second_nucs, []) self.context2pos[second_nucs].append(second_pos) self.pos2context[first_pos] = first_nucs self.pos2context[second_pos] = second_nucs # sequence context for three prime splice site for i, three_ss in enumerate(gene_seq.three_prime_seq): first_nucs = three_ss[1-index_context:1+1] second_nucs = three_ss[2-index_context:2+1] first_pos = 2*i + gene_len + five_ss_len second_pos = 2*i + gene_len + five_ss_len + 1 self.context2pos.setdefault(first_nucs, []) self.context2pos[first_nucs].append(first_pos) self.context2pos.setdefault(second_nucs, []) self.context2pos[second_nucs].append(second_pos) self.pos2context[first_pos] = first_nucs self.pos2context[second_pos] = second_nucs # hack solution for context for first nuc if gene_seq.exon_seq and gene_seq.nuc_context > 1: self.pos2context[0] = gene_seq.exon_seq[0] * 2 self.context2pos.setdefault(gene_seq.exon_seq[0]*2, []) self.context2pos[gene_seq.exon_seq[0]*2].append(0) elif gene_seq.nuc_context in [1.5, 3]: # use the nucleotide context from chasm if nuc # context is 1.5 otherwise always use a three # nucleotide context ncontext = gene_seq.nuc_context for i in range(1, len(gene_seq.exon_seq)-1): nucs = gene_seq.exon_seq[i-1:i+2] if ncontext == 1.5: context = prob2020.python.mutation_context.get_chasm_context(nucs) else: context = nucs self.context2pos.setdefault(context, []) self.context2pos[context].append(i) self.pos2context[i] = context # sequence context for five prime splice site for i, five_ss in enumerate(gene_seq.five_prime_seq): first_nucs = five_ss[:3] second_nucs = five_ss[1:4] first_pos = 2*i + gene_len second_pos = 2*i + gene_len + 1 if ncontext == 1.5: first_context = prob2020.python.mutation_context.get_chasm_context(first_nucs) second_context = prob2020.python.mutation_context.get_chasm_context(second_nucs) else: first_context = first_nucs second_context = second_nucs self.context2pos.setdefault(first_context, []) self.context2pos[first_context].append(first_pos) self.context2pos.setdefault(second_context, []) self.context2pos[second_context].append(second_pos) self.pos2context[first_pos] = first_context self.pos2context[second_pos] = second_context # sequence context for three prime splice site for i, three_ss in enumerate(gene_seq.three_prime_seq): first_nucs = three_ss[:3] second_nucs = three_ss[1:4] first_pos = 2*i + gene_len + five_ss_len second_pos = 2*i + gene_len + five_ss_len + 1 if ncontext == 1.5: first_context = prob2020.python.mutation_context.get_chasm_context(first_nucs) second_context = prob2020.python.mutation_context.get_chasm_context(second_nucs) else: first_context = first_nucs second_context = second_nucs self.context2pos.setdefault(first_context, []) self.context2pos[first_context].append(first_pos) self.context2pos.setdefault(second_context, []) self.context2pos[second_context].append(second_pos) self.pos2context[first_pos] = first_context self.pos2context[second_pos] = second_context # hack solution for context for first nuc if gene_seq.exon_seq: first_nuc = gene_seq.exon_seq[0] + gene_seq.exon_seq[:2] if ncontext == 1.5: first_context = prob2020.python.mutation_context.get_chasm_context(first_nuc) else: first_context = first_nuc self.pos2context[0] = first_context self.context2pos.setdefault(first_context, []) self.context2pos[first_context].append(0) last_nuc = gene_seq.exon_seq[-2:] + gene_seq.exon_seq[-1] if ncontext == 1.5: last_context = prob2020.python.mutation_context.get_chasm_context(last_nuc) else: last_context = last_nuc last_pos = len(gene_seq.exon_seq) - 1 self.pos2context[last_pos] = first_context self.context2pos.setdefault(last_context, []) self.context2pos[last_context].append(last_pos) else: # case where there is no context, # mutations occur with uniform probability at each # position for i in range(gene_len + five_ss_len + three_ss_len): self.pos2context[i] = 'None' self.context2pos['None'] = range(gene_len + five_ss_len + three_ss_len)
Initializes attributes defining mutation contexts and their position. The self.context2pos and self.pos2context dictionaries map from sequence context to sequence position and sequence position to sequence context, respectively. These attributes allow for randomly sampling of mutation positions while respecting sequence context in the randomization-based test. Parameters ---------- gene_seq : GeneSequence GeneSequence object from the gene_sequence module
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def random_context_pos(self, num, num_permutations, context): """Samples with replacement available positions matching the sequence context. Note: this method does random sampling only for an individual sequence context. Parameters ---------- num : int Number of positions to sample for each permutation. This is the number of actually observed mutations having the matching sequence context for this gene. num_permutations : int Number of permutations for permutation test. context : str Sequence context. Returns ------- random_pos : np.array num_permutations X num sized array that represents the randomly sampled positions for a specific context. """ # make sure provide context is valid if not self.is_valid_context(context): error_msg = 'Context ({0}) was never seen in sequence.'.format(context) raise ValueError(error_msg) # make sure sampling is a positive integer if num < 1: error_msg = ('There must be at least one sample (specified {0}) ' 'for a context'.format(num)) raise ValueError(error_msg) # randomly select from available positions that fit the specified context available_pos = self.context2pos[context] random_pos = self.prng_dict[context].choice(available_pos, (num_permutations, num)) return random_pos
Samples with replacement available positions matching the sequence context. Note: this method does random sampling only for an individual sequence context. Parameters ---------- num : int Number of positions to sample for each permutation. This is the number of actually observed mutations having the matching sequence context for this gene. num_permutations : int Number of permutations for permutation test. context : str Sequence context. Returns ------- random_pos : np.array num_permutations X num sized array that represents the randomly sampled positions for a specific context.
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def random_pos(self, context_iterable, num_permutations): """Obtains random positions w/ replacement which match sequence context. Parameters ---------- context_iterable: iterable containing two element tuple Records number of mutations in each context. context_iterable should be something like [('AA', 5), ...]. num_permutations : int Number of permutations used in the permutation test. Returns ------- position_list : list Contains context string and the randomly chosen positions for that context. """ position_list = [] for contxt, n in context_iterable: pos_array = self.random_context_pos(n, num_permutations, contxt) position_list.append([contxt, pos_array]) return position_list
Obtains random positions w/ replacement which match sequence context. Parameters ---------- context_iterable: iterable containing two element tuple Records number of mutations in each context. context_iterable should be something like [('AA', 5), ...]. num_permutations : int Number of permutations used in the permutation test. Returns ------- position_list : list Contains context string and the randomly chosen positions for that context.
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def multiprocess_permutation(bed_dict, mut_df, opts, indel_df=None): """Handles parallelization of permutations by splitting work by chromosome. """ chroms = sorted(bed_dict.keys(), key=lambda x: len(bed_dict[x]), reverse=True) multiprocess_flag = opts['processes']>0 if multiprocess_flag: num_processes = opts['processes'] else: num_processes = 1 #file_handle = open(opts['output'], 'w') file_handle = opts['handle'] mywriter = csv.writer(file_handle, delimiter='\t', lineterminator='\n') if opts['maf'] and opts['num_iterations']: header = ['Gene', 'strand', 'Chromosome', 'Start_Position', 'End_Position', 'Reference_Allele', 'Tumor_Allele', 'Context', 'DNA_Change', 'Protein_Change', 'Variant_Classification'] elif opts['maf']: header = ['Gene', 'strand', 'Chromosome', 'Start_Position', 'End_Position', 'Reference_Allele', 'Tumor_Allele', 'DNA_Change', 'Protein_Change', 'Variant_Classification', 'Tumor_Sample', 'Tumor_Type'] else: header = ['Gene', 'ID', 'gene length', 'non-silent snv', 'silent snv', 'nonsense', 'lost stop', 'splice site', 'lost start', 'missense', 'recurrent missense', 'normalized missense position entropy',] # add column header for scores, is user provided one if opts['score_dir']: header += ['Total Missense MGAEntropy', 'Total Missense VEST Score'] # add indel columns header += ['frameshift indel', 'inframe indel', 'normalized mutation entropy'] mywriter.writerow(header) num_iterations = opts['num_iterations'] # simulate indel counts if opts['summary'] and num_iterations: fs_cts, inframe_cts, gene_names = indel.simulate_indel_counts(indel_df, bed_dict, num_iterations, opts['seed']) name2ix = {gene_names[z]: z for z in range(len(gene_names))} # just count observed indels elif opts['summary']: # get gene names gene_names = [mybed.gene_name for chrom in bed_dict for mybed in bed_dict[chrom]] name2ix = {gene_names[z]: z for z in range(len(gene_names))} # initiate count vectors inframe_cts = np.zeros((1, len(gene_names))) fs_cts = np.zeros((1, len(gene_names))) # populate observed counts indel_cts_dict = indel_df['Gene'].value_counts().to_dict() fs_cts_dict = indel_df[indel.is_frameshift_annotation(indel_df)]['Gene'].value_counts().to_dict() for mygene in indel_cts_dict: if mygene in name2ix: # gene should be found in BED file annotation ix = name2ix[mygene] fs_cts[0, ix] = 0 if mygene not in fs_cts_dict else fs_cts_dict[mygene] inframe_cts[0, ix] = indel_cts_dict[mygene] - fs_cts[0, ix] # simulate snvs obs_result = [] for i in range(0, len(chroms), num_processes): if multiprocess_flag: pool = Pool(processes=num_processes) tmp_num_proc = len(chroms) - i if i + num_processes > len(chroms) else num_processes info_repeat = ((bed_dict[chroms[tmp_ix]], mut_df, opts) for tmp_ix in range(i, i+tmp_num_proc)) process_results = pool.imap(singleprocess_permutation, info_repeat) process_results.next = utils.keyboard_exit_wrapper(process_results.next) try: # iterate through each chromosome result for chrom_result in process_results: # add columns for indels if opts['summary']: tmp_chrom_result = [] for gname, grp in it.groupby(chrom_result, lambda x: x[0]): for l, row in enumerate(grp): gene_ix = name2ix[gname] fs_count = fs_cts[l, gene_ix] inframe_count = inframe_cts[l, gene_ix] missense_pos_ct = list(row.pop(-1).values()) # missense codon counts silent_pos_ct = [1 for l in range(row[4])] inactivating_ct = sum(row[5:9]) + fs_count tmp_count_list = missense_pos_ct + silent_pos_ct + [inactivating_ct, inframe_count] norm_ent = math.normalized_mutation_entropy(tmp_count_list) tmp_chrom_result.append(row+[fs_count, inframe_count, norm_ent]) chrom_result = tmp_chrom_result # write output to file mywriter.writerows(chrom_result) except KeyboardInterrupt: pool.close() pool.join() logger.info('Exited by user. ctrl-c') sys.exit(0) pool.close() pool.join() else: # perform simulation info = (bed_dict[chroms[i]], mut_df, opts) chrom_results = singleprocess_permutation(info) # add indel columns if opts['summary']: tmp_chrom_result = [] for gname, grp in it.groupby(chrom_results, lambda x: x[0]): for l, row in enumerate(grp): gene_ix = name2ix[gname] fs_count = fs_cts[l, gene_ix] inframe_count = inframe_cts[l, gene_ix] missense_pos_ct = list(row.pop(-1).values()) # missense codon counts silent_pos_ct = [1 for l in range(row[4])] inactivating_ct = sum(row[5:9]) + fs_count tmp_count_list = missense_pos_ct + silent_pos_ct + [inactivating_ct, inframe_count] norm_ent = math.normalized_mutation_entropy(tmp_count_list) tmp_chrom_result.append(row+[fs_count, inframe_count, norm_ent]) chrom_results = tmp_chrom_result # write to file mywriter.writerows(chrom_results)
Handles parallelization of permutations by splitting work by chromosome.
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def multiprocess_permutation(bed_dict, mut_df, opts): """Handles parallelization of permutations by splitting work by chromosome. """ chroms = sorted(bed_dict.keys()) multiprocess_flag = opts['processes']>0 if multiprocess_flag: num_processes = opts['processes'] else: num_processes = 1 num_permutations = opts['num_permutations'] if not opts['by_sample']: obs_result = [] else: uniq_samp = mut_df['Tumor_Sample'].unique() obs_result = pd.DataFrame(np.zeros((len(uniq_samp), len(cols))), index=uniq_samp, columns=cols) # initialize list containing output if not opts['score_dir']: result_list = [[0, 0, 0, 0, 0, 0, 0] for k in range(num_permutations)] else: result_list = [[0, 0, 0, 0, 0, 0, 0, 0, 0] for k in range(num_permutations)] # iterate over each chromosome for i in range(0, len(chroms), num_processes): if multiprocess_flag: pool = Pool(processes=num_processes) tmp_num_proc = len(chroms) - i if i + num_processes > len(chroms) else num_processes info_repeat = ((bed_dict[chroms[tmp_ix]], mut_df, opts) for tmp_ix in range(i, i+tmp_num_proc)) process_results = pool.imap(singleprocess_permutation, info_repeat) process_results.next = utils.keyboard_exit_wrapper(process_results.next) try: for chrom_result, obs_mutations in process_results: for j in range(num_permutations): result_list[j][0] += chrom_result[j][0] result_list[j][1] += chrom_result[j][1] result_list[j][2] += chrom_result[j][2] result_list[j][3] += chrom_result[j][3] result_list[j][4] += chrom_result[j][4] result_list[j][5] += chrom_result[j][5] result_list[j][6] += chrom_result[j][6] if opts['score_dir']: result_list[j][7] += chrom_result[j][7] result_list[j][8] += chrom_result[j][8] if not opts['by_sample']: obs_result.append(obs_mutations) else: obs_result = obs_result + obs_mutations except KeyboardInterrupt: pool.close() pool.join() logger.info('Exited by user. ctrl-c') sys.exit(0) pool.close() pool.join() else: info = (bed_dict[chroms[i]], mut_df, opts) chrom_result, obs_mutations = singleprocess_permutation(info) for j in range(num_permutations): result_list[j][0] += chrom_result[j][0] result_list[j][1] += chrom_result[j][1] result_list[j][2] += chrom_result[j][2] result_list[j][3] += chrom_result[j][3] result_list[j][4] += chrom_result[j][4] result_list[j][5] += chrom_result[j][5] result_list[j][6] += chrom_result[j][6] if opts['score_dir']: result_list[j][7] += chrom_result[j][7] result_list[j][8] += chrom_result[j][8] if not opts['by_sample']: obs_result.append(obs_mutations) else: obs_result = obs_result + obs_mutations return result_list, obs_result
Handles parallelization of permutations by splitting work by chromosome.
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def retrieve_scores(gname, sdir, codon_pos, germ_aa, somatic_aa, default_mga=5., default_vest=0, no_file_flag=-1): """Retrieves scores from pickle files. Used by summary script. """ # get variant types #var_class = cutils.get_variant_classification(germ_aa, somatic_aa, codon_pos) # get information about MGA entropy mga_path = os.path.join(sdir, gname+".mgaentropy.pickle") if os.path.exists(mga_path): if sys.version_info < (3,): # python 2.7 way with open(mga_path) as handle: mga_ent = pickle.load(handle) else: # python 3.X way with open(mga_path, 'rb') as handle: mga_ent = pickle.load(handle, encoding='latin-1') else: mga_ent = None missense_pos = [p for i, p in enumerate(codon_pos) if (germ_aa[i]!=somatic_aa[i]) and (germ_aa[i] not in ['-', '*', 'Splice_Site']) and (somatic_aa[i] not in ['-', '*', 'Splice_Site'])] total_mga_ent = compute_mga_entropy_stat(mga_ent, missense_pos, sum, default_mga) #mga_ent_ixs = [codon_pos[i] for i in range(len(var_class)) #if var_class[i] == 'Missense_Mutation'] #len_mga_ent = len(mga_ent) #mga_ent_scores = [mga_ent[ix] for ix in mga_ent_ixs if ix < len_mga_ent] #if mga_ent_scores: #total_mga_ent = sum(mga_ent_scores) #else: #total_mga_ent = default_mga #else: #total_mga_ent = no_file_flag # get information about VEST scores vest_path = os.path.join(sdir, gname+".vest.pickle") if os.path.exists(vest_path): if sys.version_info < (3,): # python 2.7 way with open(vest_path) as handle: vest_score = pickle.load(handle) else: # python 3.X way with open(vest_path, 'rb') as handle: vest_score = pickle.load(handle, encoding='latin-1') else: vest_score = None total_vest = compute_vest_stat(vest_score, germ_aa, somatic_aa, codon_pos, stat_func=sum, default_val=default_vest) #vest_scores = [vest_score.get(codon_pos[i]+1, {}).get(germ_aa[i], {}).get(somatic_aa[i], default_vest) #for i in range(len(var_class)) #if var_class[i] == 'Missense_Mutation'] #total_vest = sum(vest_scores) #else: #total_vest = no_file_flag return total_mga_ent, total_vest
Retrieves scores from pickle files. Used by summary script.
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def read_vest_pickle(gname, score_dir): """Read in VEST scores for given gene. Parameters ---------- gname : str name of gene score_dir : str directory containing vest scores Returns ------- gene_vest : dict or None dict containing vest scores for gene. Returns None if not found. """ vest_path = os.path.join(score_dir, gname+".vest.pickle") if os.path.exists(vest_path): if sys.version_info < (3,): with open(vest_path) as handle: gene_vest = pickle.load(handle) else: with open(vest_path, 'rb') as handle: gene_vest = pickle.load(handle, encoding='latin-1') return gene_vest else: return None
Read in VEST scores for given gene. Parameters ---------- gname : str name of gene score_dir : str directory containing vest scores Returns ------- gene_vest : dict or None dict containing vest scores for gene. Returns None if not found.
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def compute_vest_stat(vest_dict, ref_aa, somatic_aa, codon_pos, stat_func=np.mean, default_val=0.0): """Compute missense VEST score statistic. Note: non-missense mutations are intentially not filtered out and will take a default value of zero. Parameters ---------- vest_dict : dict dictionary containing vest scores across the gene of interest ref_aa: list of str list of reference amino acids somatic_aa: list of str somatic mutation aa codon_pos : list of int position of codon in protein sequence stat_func : function, default=np.mean function that calculates a statistic default_val : float default value to return if there are no mutations Returns ------- score_stat : float vest score statistic for provided mutation list """ # return default value if VEST scores are missing if vest_dict is None: return default_val # fetch scores myscores = fetch_vest_scores(vest_dict, ref_aa, somatic_aa, codon_pos) # calculate mean score if myscores: score_stat = stat_func(myscores) else: score_stat = default_val return score_stat
Compute missense VEST score statistic. Note: non-missense mutations are intentially not filtered out and will take a default value of zero. Parameters ---------- vest_dict : dict dictionary containing vest scores across the gene of interest ref_aa: list of str list of reference amino acids somatic_aa: list of str somatic mutation aa codon_pos : list of int position of codon in protein sequence stat_func : function, default=np.mean function that calculates a statistic default_val : float default value to return if there are no mutations Returns ------- score_stat : float vest score statistic for provided mutation list
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def compute_mga_entropy_stat(mga_vec, codon_pos, stat_func=np.mean, default_val=0.0): """Compute MGA entropy conservation statistic Parameters ---------- mga_vec : np.array numpy vector containing MGA Entropy conservation scores for residues codon_pos : list of int position of codon in protein sequence stat_func : function, default=np.mean function that calculates a statistic default_val : float default value to return if there are no mutations Returns ------- score_stat : float MGA entropy score statistic for provided mutation list """ # return default value if VEST scores are missing if mga_vec is None: return default_val # fetch scores myscores = fetch_mga_scores(mga_vec, codon_pos) # calculate mean score if myscores is not None and len(myscores): score_stat = stat_func(myscores) else: score_stat = default_val return score_stat
Compute MGA entropy conservation statistic Parameters ---------- mga_vec : np.array numpy vector containing MGA Entropy conservation scores for residues codon_pos : list of int position of codon in protein sequence stat_func : function, default=np.mean function that calculates a statistic default_val : float default value to return if there are no mutations Returns ------- score_stat : float MGA entropy score statistic for provided mutation list
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def fetch_vest_scores(vest_dict, ref_aa, somatic_aa, codon_pos, default_vest=0.0): """Get VEST scores from pre-computed scores in dictionary. Note: either all mutations should be missense or non-missense intended to have value equal to default. Parameters ---------- vest_dict : dict dictionary containing vest scores across the gene of interest ref_aa: list of str list of reference amino acids somatic_aa: list of str somatic mutation aa codon_pos: list of int position of codon in protein sequence default_vest: float, default=0.0 value to use if VEST score not available for a given mutation Returns ------- vest_score_list: list score results for mutations """ vest_score_list = [] for i in range(len(somatic_aa)): # make sure position is valid if codon_pos[i] is not None: tmp_score = vest_dict.get(codon_pos[i]+1, {}).get(ref_aa[i], {}).get(somatic_aa[i], default_vest) else: tmp_score = 0.0 vest_score_list.append(tmp_score) return vest_score_list
Get VEST scores from pre-computed scores in dictionary. Note: either all mutations should be missense or non-missense intended to have value equal to default. Parameters ---------- vest_dict : dict dictionary containing vest scores across the gene of interest ref_aa: list of str list of reference amino acids somatic_aa: list of str somatic mutation aa codon_pos: list of int position of codon in protein sequence default_vest: float, default=0.0 value to use if VEST score not available for a given mutation Returns ------- vest_score_list: list score results for mutations
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def fetch_mga_scores(mga_vec, codon_pos, default_mga=None): """Get MGAEntropy scores from pre-computed scores in array. Parameters ---------- mga_vec : np.array numpy vector containing MGA Entropy conservation scores for residues codon_pos: list of int position of codon in protein sequence default_mga: float or None, default=None value to use if MGA entropy score not available for a given mutation. Drop mutations if no default specified. Returns ------- mga_ent_scores : np.array score results for MGA entropy conservation """ # keep only positions in range of MGAEntropy scores len_mga = len(mga_vec) good_codon_pos = [p for p in codon_pos if p < len_mga] # get MGAEntropy scores if good_codon_pos: mga_ent_scores = mga_vec[good_codon_pos] else: mga_ent_scores = None return mga_ent_scores
Get MGAEntropy scores from pre-computed scores in array. Parameters ---------- mga_vec : np.array numpy vector containing MGA Entropy conservation scores for residues codon_pos: list of int position of codon in protein sequence default_mga: float or None, default=None value to use if MGA entropy score not available for a given mutation. Drop mutations if no default specified. Returns ------- mga_ent_scores : np.array score results for MGA entropy conservation
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def read_neighbor_graph_pickle(gname, graph_dir): """Read in neighbor graph for given gene. Parameters ---------- gname : str name of gene graph_dir : str directory containing gene graphs Returns ------- gene_graph : dict or None neighbor graph as dict for gene. Returns None if not found. """ graph_path = os.path.join(graph_dir, gname+".pickle") if os.path.exists(graph_path): with open(graph_path) as handle: gene_graph = pickle.load(handle) return gene_graph else: return None
Read in neighbor graph for given gene. Parameters ---------- gname : str name of gene graph_dir : str directory containing gene graphs Returns ------- gene_graph : dict or None neighbor graph as dict for gene. Returns None if not found.
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def compute_ng_stat(gene_graph, pos_ct, alpha=.5): """Compute the clustering score for the gene on its neighbor graph. Parameters ---------- gene_graph : dict Graph of spatially near codons. keys = nodes, edges = key -> value. pos_ct : dict missense mutation count for each codon alpha : float smoothing factor Returns ------- graph_score : float score measuring the clustering of missense mutations in the graph coverage : int number of nodes that received non-zero weight """ # skip if there are no missense mutations if not len(pos_ct): return 1.0, 0 max_pos = max(gene_graph) codon_vals = np.zeros(max_pos+1) # smooth out mutation counts for pos in pos_ct: mut_count = pos_ct[pos] # update neighbor values neighbors = list(gene_graph[pos]) num_neighbors = len(neighbors) codon_vals[neighbors] += alpha*mut_count # update self-value codon_vals[pos] += (1-alpha)*mut_count # compute the normalized entropy #total_cts = float(np.count_nonzero(codon_vals)) #graph_score = mymath.normalized_mutation_entropy(codon_vals, total_cts=total_cts) # compute regular entropy p = codon_vals / np.sum(codon_vals) graph_score = mymath.shannon_entropy(p) # get coverage coverage = np.count_nonzero(p) return graph_score, coverage
Compute the clustering score for the gene on its neighbor graph. Parameters ---------- gene_graph : dict Graph of spatially near codons. keys = nodes, edges = key -> value. pos_ct : dict missense mutation count for each codon alpha : float smoothing factor Returns ------- graph_score : float score measuring the clustering of missense mutations in the graph coverage : int number of nodes that received non-zero weight
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def count_frameshift_total(mut_df, bed_path, use_unmapped=False, to_zero_based=False): """Count frameshifts for each gene. Parameters ---------- mut_df : pd.DataFrame mutation input bed_path : str path to BED file containing reference tx for genes use_unmapped : Bool flag indicating whether to include frameshifts not mapping to reference tx to_zero_based : Bool whether to convert end-coordinate to zero based for analysis Returns ------- fs_cts_df : pd.DataFrame contains both total frameshift counts and frameshift counts not mappable to the reference transcript. """ if to_zero_based: mut_df['Start_Position'] = mut_df['Start_Position'] - 1 fs_cts = {} # frameshift count information for each gene fs_df = indel.keep_frameshifts(mut_df) for bed in utils.bed_generator(bed_path): gene_df = fs_df[fs_df['Gene']==bed.gene_name] # find it frameshift actually is on gene annotation fs_pos = [] for ix, row in gene_df.iterrows(): indel_pos = [row['Start_Position'], row['End_Position']] coding_pos = bed.query_position(bed.strand, row['Chromosome'], indel_pos) fs_pos.append(coding_pos) # mark frameshifts that could not be mapped to reference tx gene_df['unmapped'] = [(1 if x is None else 0) for x in fs_pos] total_fs = len(gene_df) unmapped_fs = len(gene_df[gene_df['unmapped']==1]) # filter out frameshifts that did not match reference tx if not use_unmapped: gene_df = gene_df[gene_df['unmapped']==0] total_fs -= unmapped_fs info = [total_fs, unmapped_fs,] fs_cts[bed.gene_name] = info # prepare counts into a dataframe fs_cts_df = pd.DataFrame.from_dict(fs_cts, orient='index') cols = ['total', 'unmapped',] fs_cts_df.columns = cols return fs_cts_df
Count frameshifts for each gene. Parameters ---------- mut_df : pd.DataFrame mutation input bed_path : str path to BED file containing reference tx for genes use_unmapped : Bool flag indicating whether to include frameshifts not mapping to reference tx to_zero_based : Bool whether to convert end-coordinate to zero based for analysis Returns ------- fs_cts_df : pd.DataFrame contains both total frameshift counts and frameshift counts not mappable to the reference transcript.
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def _fetch_3ss_fasta(fasta, gene_name, exon_num, chrom, strand, start, end): """Retreives the 3' SS sequence flanking the specified exon. Returns a string in fasta format with the first line containing a ">" and the second line contains the two base pairs of 3' SS. Parameters ---------- fasta : pysam.Fastafile fasta object from pysam gene_name : str gene name used for fasta seq id exon_num : int the `exon_num` exon, used for seq id chrom : str chromsome strand : str strand, {'+', '-'} start : int 0-based start position end : int 0-based end position Returns ------- ss_fasta : str string in fasta format with first line being seq id """ if strand == '-': ss_seq = fasta.fetch(reference=chrom, start=end-1, end=end+3) ss_seq = utils.rev_comp(ss_seq) elif strand == '+': ss_seq = fasta.fetch(reference=chrom, start=start-3, end=start+1) ss_fasta = '>{0};exon{1};3SS\n{2}\n'.format(gene_name, exon_num, ss_seq.upper()) return ss_fasta
Retreives the 3' SS sequence flanking the specified exon. Returns a string in fasta format with the first line containing a ">" and the second line contains the two base pairs of 3' SS. Parameters ---------- fasta : pysam.Fastafile fasta object from pysam gene_name : str gene name used for fasta seq id exon_num : int the `exon_num` exon, used for seq id chrom : str chromsome strand : str strand, {'+', '-'} start : int 0-based start position end : int 0-based end position Returns ------- ss_fasta : str string in fasta format with first line being seq id
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def fetch_gene_fasta(gene_bed, fasta_obj): """Retreive gene sequences in FASTA format. Parameters ---------- gene_bed : BedLine BedLine object representing a single gene fasta_obj : pysam.Fastafile fasta object for index retreival of sequence Returns ------- gene_fasta : str sequence of gene in FASTA format """ gene_fasta = '' strand = gene_bed.strand exons = gene_bed.get_exons() if strand == '-': exons.reverse() # order exons 5' to 3', so reverse if '-' strand # iterate over exons for i, exon in enumerate(exons): exon_seq = fasta_obj.fetch(reference=gene_bed.chrom, start=exon[0], end=exon[1]).upper() if strand == '-': exon_seq = utils.rev_comp(exon_seq) exon_fasta = '>{0};exon{1}\n{2}\n'.format(gene_bed.gene_name, i, exon_seq) # get splice site sequence if len(exons) == 1: # splice sites don't matter if there is no splicing ss_fasta = '' elif i == 0: # first exon only, get 3' SS ss_fasta = _fetch_5ss_fasta(fasta_obj, gene_bed.gene_name, i, gene_bed.chrom, strand, exon[0], exon[1]) elif i == (len(exons) - 1): # last exon only, get 5' SS ss_fasta = _fetch_3ss_fasta(fasta_obj, gene_bed.gene_name, i, gene_bed.chrom, strand, exon[0], exon[1]) else: # middle exon, get bot 5' and 3' SS fasta_3ss = _fetch_3ss_fasta(fasta_obj, gene_bed.gene_name, i, gene_bed.chrom, strand, exon[0], exon[1]) fasta_5ss = _fetch_5ss_fasta(fasta_obj, gene_bed.gene_name, i, gene_bed.chrom, strand, exon[0], exon[1]) ss_fasta = fasta_5ss + fasta_3ss gene_fasta += exon_fasta + ss_fasta return gene_fasta
Retreive gene sequences in FASTA format. Parameters ---------- gene_bed : BedLine BedLine object representing a single gene fasta_obj : pysam.Fastafile fasta object for index retreival of sequence Returns ------- gene_fasta : str sequence of gene in FASTA format
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def _reset_seq(self): """Updates attributes for gene represented in the self.bed attribute. Sequences are always upper case. """ exon_seq_list, five_ss_seq_list, three_ss_seq_list = self._fetch_seq() self.exon_seq = ''.join(exon_seq_list) self.three_prime_seq = three_ss_seq_list self.five_prime_seq = five_ss_seq_list self._to_upper()
Updates attributes for gene represented in the self.bed attribute. Sequences are always upper case.
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def add_germline_variants(self, germline_nucs, coding_pos): """Add potential germline variants into the nucleotide sequence. Sequenced individuals may potentially have a SNP at a somatic mutation position. Therefore they may differ from the reference genome. This method updates the gene germline gene sequence to match the actual individual. Parameters ---------- germline_nucs : list of str list of DNA nucleotides containing the germline letter coding_pos : int 0-based nucleotide position in coding sequence NOTE: the self.exon_seq attribute is updated, no return value """ if len(germline_nucs) != len(coding_pos): raise ValueError('Each germline nucleotide should have a coding position') es = list(self.exon_seq) for i in range(len(germline_nucs)): gl_nuc, cpos = germline_nucs[i].upper(), coding_pos[i] if not utils.is_valid_nuc(gl_nuc): raise ValueError('{0} is not a valid nucleotide'.format(gl_nuc)) if cpos >= 0: es[cpos] = gl_nuc self.exon_seq = ''.join(es)
Add potential germline variants into the nucleotide sequence. Sequenced individuals may potentially have a SNP at a somatic mutation position. Therefore they may differ from the reference genome. This method updates the gene germline gene sequence to match the actual individual. Parameters ---------- germline_nucs : list of str list of DNA nucleotides containing the germline letter coding_pos : int 0-based nucleotide position in coding sequence NOTE: the self.exon_seq attribute is updated, no return value
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def _to_upper(self): """Convert sequences to upper case.""" self.exon_seq = self.exon_seq.upper() self.three_prime_seq = [s.upper() for s in self.three_prime_seq] self.five_prime_seq = [s.upper() for s in self.five_prime_seq]
Convert sequences to upper case.
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def _fetch_seq(self): """Fetches gene sequence from PySAM fasta object. Returns ------- exons : list of str list of exon nucleotide sequences five_prime_ss : list of str list of 5' splice site sequences three_prime_ss : list of str list of 3' splice site sequences """ exons = [] three_prime_ss = [] five_prime_ss = [] num_exons = self.bed.get_num_exons() for i in range(num_exons): # add exon sequence tmp_id = '{0};exon{1}'.format(self.bed.gene_name, i) tmp_exon = self.fasta.fetch(reference=tmp_id) exons.append(tmp_exon) # add splice site sequence tmp_id_3ss = '{0};3SS'.format(tmp_id) tmp_id_5ss = '{0};5SS'.format(tmp_id) if num_exons == 1: pass elif i == 0: tmp_5ss = self.fasta.fetch(tmp_id_5ss) five_prime_ss.append(tmp_5ss) elif i == (num_exons - 1): tmp_3ss = self.fasta.fetch(tmp_id_3ss) three_prime_ss.append(tmp_3ss) else: tmp_3ss = self.fasta.fetch(tmp_id_3ss) tmp_5ss = self.fasta.fetch(tmp_id_5ss) three_prime_ss.append(tmp_3ss) five_prime_ss.append(tmp_5ss) return exons, five_prime_ss, three_prime_ss
Fetches gene sequence from PySAM fasta object. Returns ------- exons : list of str list of exon nucleotide sequences five_prime_ss : list of str list of 5' splice site sequences three_prime_ss : list of str list of 3' splice site sequences
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def correct_chrom_names(chroms): """Make sure chromosome names follow UCSC chr convention.""" chrom_list = [] for chrom in chroms: # fix chrom numbering chrom = str(chrom) chrom = chrom.replace('23', 'X') chrom = chrom.replace('24', 'Y') chrom = chrom.replace('25', 'Mt') if not chrom.startswith('chr'): chrom = 'chr' + chrom chrom_list.append(chrom) return chrom_list
Make sure chromosome names follow UCSC chr convention.
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def fishers_method(pvals): """Fisher's method for combining independent p-values.""" pvals = np.asarray(pvals) degrees_of_freedom = 2 * pvals.size chisq_stat = np.sum(-2*np.log(pvals)) fishers_pval = stats.chi2.sf(chisq_stat, degrees_of_freedom) return fishers_pval
Fisher's method for combining independent p-values.
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def cummin(x): """A python implementation of the cummin function in R""" for i in range(1, len(x)): if x[i-1] < x[i]: x[i] = x[i-1] return x
A python implementation of the cummin function in R
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def bh_fdr(pval): """A python implementation of the Benjamani-Hochberg FDR method. This code should always give precisely the same answer as using p.adjust(pval, method="BH") in R. Parameters ---------- pval : list or array list/array of p-values Returns ------- pval_adj : np.array adjusted p-values according the benjamani-hochberg method """ pval_array = np.array(pval) sorted_order = np.argsort(pval_array) original_order = np.argsort(sorted_order) pval_array = pval_array[sorted_order] # calculate the needed alpha n = float(len(pval)) pval_adj = np.zeros(int(n)) i = np.arange(1, int(n)+1, dtype=float)[::-1] # largest to smallest pval_adj = np.minimum(1, cummin(n/i * pval_array[::-1]))[::-1] return pval_adj[original_order]
A python implementation of the Benjamani-Hochberg FDR method. This code should always give precisely the same answer as using p.adjust(pval, method="BH") in R. Parameters ---------- pval : list or array list/array of p-values Returns ------- pval_adj : np.array adjusted p-values according the benjamani-hochberg method
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def calc_deleterious_p_value(mut_info, unmapped_mut_info, sc, gs, bed, num_permutations, stop_thresh, del_threshold, pseudo_count, seed=None): """Calculates the p-value for the number of inactivating SNV mutations. Calculates p-value based on how many simulations exceed the observed value. Parameters ---------- mut_info : dict contains codon and amino acid residue information for mutations mappable to provided reference tx. unmapped_mut_info : dict contains codon/amino acid residue info for mutations that are NOT mappable to provided reference tx. fs_ct : int number of frameshifts for gene prob_inactive : float proportion of inactivating mutations out of total over all genes sc : SequenceContext object contains the nucleotide contexts for a gene such that new random positions can be obtained while respecting nucleotide context. gs : GeneSequence contains gene sequence bed : BedLine just used to return gene name num_permutations : int number of permutations to perform to estimate p-value. more permutations means more precision on the p-value. seed : int (Default: None) seed number to random number generator (None to be randomly set) """ #prng = np.random.RandomState(seed) if len(mut_info) > 0: mut_info['Coding Position'] = mut_info['Coding Position'].astype(int) mut_info['Context'] = mut_info['Coding Position'].apply(lambda x: sc.pos2context[x]) # group mutations by context cols = ['Context', 'Tumor_Allele'] unmapped_mut_df = pd.DataFrame(unmapped_mut_info) tmp_df = pd.concat([mut_info[cols], unmapped_mut_df[cols]]) context_cts = tmp_df['Context'].value_counts() context_to_mutations = dict((name, group['Tumor_Allele']) for name, group in tmp_df.groupby('Context')) # get deleterious info for actual mutations aa_mut_info = mc.get_aa_mut_info(mut_info['Coding Position'], mut_info['Tumor_Allele'].tolist(), gs) ref_aa = aa_mut_info['Reference AA'] + unmapped_mut_info['Reference AA'] somatic_aa = aa_mut_info['Somatic AA'] + unmapped_mut_info['Somatic AA'] codon_pos = aa_mut_info['Codon Pos'] + unmapped_mut_info['Codon Pos'] num_del = cutils.calc_deleterious_info(ref_aa, somatic_aa, codon_pos) #num_del = fs_ct + num_snv_del # skip permutation test if number of deleterious mutations is not at # least meet some user-specified threshold if num_del >= del_threshold: # perform permutations del_p_value = pm.deleterious_permutation(num_del, context_cts, context_to_mutations, sc, # sequence context obj gs, # gene sequence obj num_permutations, stop_thresh, pseudo_count) else: del_p_value = None else: num_del = 0 del_p_value = None result = [bed.gene_name, num_del, del_p_value] return result
Calculates the p-value for the number of inactivating SNV mutations. Calculates p-value based on how many simulations exceed the observed value. Parameters ---------- mut_info : dict contains codon and amino acid residue information for mutations mappable to provided reference tx. unmapped_mut_info : dict contains codon/amino acid residue info for mutations that are NOT mappable to provided reference tx. fs_ct : int number of frameshifts for gene prob_inactive : float proportion of inactivating mutations out of total over all genes sc : SequenceContext object contains the nucleotide contexts for a gene such that new random positions can be obtained while respecting nucleotide context. gs : GeneSequence contains gene sequence bed : BedLine just used to return gene name num_permutations : int number of permutations to perform to estimate p-value. more permutations means more precision on the p-value. seed : int (Default: None) seed number to random number generator (None to be randomly set)
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def calc_protein_p_value(mut_info, unmapped_mut_info, sc, gs, bed, graph_dir, num_permutations, stop_thresh, min_recurrent, min_fraction): """Computes the p-value for clustering on a neighbor graph composed of codons connected with edges if they are spatially near in 3D protein structure. Parameters ---------- Returns ------- """ if len(mut_info) > 0: mut_info['Coding Position'] = mut_info['Coding Position'].astype(int) mut_info['Context'] = mut_info['Coding Position'].apply(lambda x: sc.pos2context[x]) # group mutations by context cols = ['Context', 'Tumor_Allele'] unmapped_mut_df = pd.DataFrame(unmapped_mut_info) tmp_df = pd.concat([mut_info[cols], unmapped_mut_df[cols]]) context_cts = tmp_df['Context'].value_counts() context_to_mutations = dict((name, group['Tumor_Allele']) for name, group in tmp_df.groupby('Context')) # get vest scores for gene if directory provided if graph_dir: gene_graph = scores.read_neighbor_graph_pickle(bed.gene_name, graph_dir) if gene_graph is None: logger.warning('Could not find neighbor graph for {0}, skipping . . .'.format(bed.gene_name)) else: gene_graph = None # get recurrent info for actual mutations aa_mut_info = mc.get_aa_mut_info(mut_info['Coding Position'], mut_info['Tumor_Allele'].tolist(), gs) codon_pos = aa_mut_info['Codon Pos'] + unmapped_mut_info['Codon Pos'] ref_aa = aa_mut_info['Reference AA'] + unmapped_mut_info['Reference AA'] somatic_aa = aa_mut_info['Somatic AA'] + unmapped_mut_info['Somatic AA'] num_recurrent, pos_ent, delta_pos_ent, pos_ct = cutils.calc_pos_info(codon_pos, ref_aa, somatic_aa, min_frac=min_fraction, min_recur=min_recurrent) try: # get vest score for actual mutations graph_score, coverage = scores.compute_ng_stat(gene_graph, pos_ct) # perform simulations to get p-value protein_p_value, norm_graph_score = pm.protein_permutation( graph_score, len(pos_ct), context_cts, context_to_mutations, sc, # sequence context obj gs, # gene sequence obj gene_graph, num_permutations, stop_thresh ) except Exception as err: exc_info = sys.exc_info() norm_graph_score = 0.0 protein_p_value = 1.0 logger.warning('Codon numbering problem with '+bed.gene_name) else: norm_graph_score = 0.0 protein_p_value = 1.0 num_recurrent = 0 result = [bed.gene_name, num_recurrent, norm_graph_score, protein_p_value] return result
Computes the p-value for clustering on a neighbor graph composed of codons connected with edges if they are spatially near in 3D protein structure. Parameters ---------- Returns -------
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def shannon_entropy(p): """Calculates shannon entropy in bits. Parameters ---------- p : np.array array of probabilities Returns ------- shannon entropy in bits """ return -np.sum(np.where(p!=0, p * np.log2(p), 0))
Calculates shannon entropy in bits. Parameters ---------- p : np.array array of probabilities Returns ------- shannon entropy in bits
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def normalized_mutation_entropy(counts, total_cts=None): """Calculate the normalized mutation entropy based on a list/array of mutation counts. Note: Any grouping of mutation counts together should be done before hand Parameters ---------- counts : np.array_like array/list of mutation counts Returns ------- norm_ent : float normalized entropy of mutation count distribution. """ cts = np.asarray(counts, dtype=float) if total_cts is None: total_cts = np.sum(cts) if total_cts > 1: p = cts / total_cts ent = shannon_entropy(p) max_ent = max_shannon_entropy(total_cts) norm_ent = ent / max_ent else: norm_ent = 1.0 return norm_ent
Calculate the normalized mutation entropy based on a list/array of mutation counts. Note: Any grouping of mutation counts together should be done before hand Parameters ---------- counts : np.array_like array/list of mutation counts Returns ------- norm_ent : float normalized entropy of mutation count distribution.
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def kl_divergence(p, q): """Compute the Kullback-Leibler (KL) divergence for discrete distributions. Parameters ---------- p : np.array "Ideal"/"true" Probability distribution q : np.array Approximation of probability distribution p Returns ------- kl : float KL divergence of approximating p with the distribution q """ # make sure numpy arrays are floats p = p.astype(float) q = q.astype(float) # compute kl divergence kl = np.sum(np.where(p!=0, p*np.log2(p/q), 0)) return kl
Compute the Kullback-Leibler (KL) divergence for discrete distributions. Parameters ---------- p : np.array "Ideal"/"true" Probability distribution q : np.array Approximation of probability distribution p Returns ------- kl : float KL divergence of approximating p with the distribution q
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def js_divergence(p, q): """Compute the Jensen-Shannon Divergence between two discrete distributions. Parameters ---------- p : np.array probability mass array (sums to 1) q : np.array probability mass array (sums to 1) Returns ------- js_div : float js divergence between the two distrubtions """ m = .5 * (p+q) js_div = .5*kl_divergence(p, m) + .5*kl_divergence(q, m) return js_div
Compute the Jensen-Shannon Divergence between two discrete distributions. Parameters ---------- p : np.array probability mass array (sums to 1) q : np.array probability mass array (sums to 1) Returns ------- js_div : float js divergence between the two distrubtions
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def js_distance(p, q): """Compute the Jensen-Shannon distance between two discrete distributions. NOTE: JS divergence is not a metric but the sqrt of JS divergence is a metric and is called the JS distance. Parameters ---------- p : np.array probability mass array (sums to 1) q : np.array probability mass array (sums to 1) Returns ------- js_dist : float Jensen-Shannon distance between two discrete distributions """ js_dist = np.sqrt(js_divergence(p, q)) return js_dist
Compute the Jensen-Shannon distance between two discrete distributions. NOTE: JS divergence is not a metric but the sqrt of JS divergence is a metric and is called the JS distance. Parameters ---------- p : np.array probability mass array (sums to 1) q : np.array probability mass array (sums to 1) Returns ------- js_dist : float Jensen-Shannon distance between two discrete distributions
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def _filter_utr(self, ex): """Filter out UTR regions from the exon list (ie retain only coding regions). Coding regions are defined by the thickStart and thickEnd attributes. Parameters ---------- ex : list of tuples list of exon positions, [(ex1_start, ex1_end), ...] Returns ------- filtered_exons : list of tuples exons with UTR regions "chopped" out """ # define coding region coding_start = int(self.bed_tuple.thickStart) coding_end = int(self.bed_tuple.thickEnd) if (coding_end - coding_start) < 3: # coding regions should have at least one codon, otherwise the # region is invalid and does not indicate an actually coding region logger.debug('{0} has an invalid coding region specified by thickStart ' 'and thickEnd (only {1} bps long). This gene is possibly either ' 'a non-coding transcript or a pseudo gene.'.format(self.gene_name, coding_end-coding_start)) return [] filtered_exons = [] for exon in ex: if exon[0] > coding_end and exon[1] > coding_end: # exon has no coding region pass elif exon[0] < coding_start and exon[1] < coding_start: # exon has no coding region pass elif exon[0] <= coding_start and exon[1] >= coding_end: # coding region entirely contained within one exon filtered_exons.append((coding_start, coding_end)) elif exon[0] <= coding_start and exon[1] < coding_end: # only beginning of exon contains UTR filtered_exons.append((coding_start, exon[1])) elif exon[0] > coding_start and exon[1] >= coding_end: # only end part of exon contains UTR filtered_exons.append((exon[0], coding_end)) elif exon[0] > coding_start and exon[1] < coding_end: # entire exon is coding filtered_exons.append(exon) else: # exon is only a UTR pass return filtered_exons
Filter out UTR regions from the exon list (ie retain only coding regions). Coding regions are defined by the thickStart and thickEnd attributes. Parameters ---------- ex : list of tuples list of exon positions, [(ex1_start, ex1_end), ...] Returns ------- filtered_exons : list of tuples exons with UTR regions "chopped" out
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def _init_exons(self): """Sets a list of position intervals for each exon. Only coding regions as defined by thickStart and thickEnd are kept. Exons are stored in the self.exons attribute. """ exon_starts = [self.chrom_start + int(s) for s in self.bed_tuple.blockStarts.strip(',').split(',')] exon_sizes = list(map(int, self.bed_tuple.blockSizes.strip(',').split(','))) # get chromosome intervals exons = [(exon_starts[i], exon_starts[i] + exon_sizes[i]) for i in range(len(exon_starts))] no_utr_exons = self._filter_utr(exons) self.exons = no_utr_exons self.exon_lens = [e[1] - e[0] for e in self.exons] self.num_exons = len(self.exons) self.cds_len = sum(self.exon_lens) self.five_ss_len = 2*(self.num_exons-1) self.three_ss_len = 2*(self.num_exons-1) self._init_splice_site_pos()
Sets a list of position intervals for each exon. Only coding regions as defined by thickStart and thickEnd are kept. Exons are stored in the self.exons attribute.
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def init_genome_coordinates(self) : """Creates the self.seqpos2genome dictionary that converts positions relative to the sequence to genome coordinates.""" self.seqpos2genome = {} # record genome positions for each sequence position seq_pos = 0 for estart, eend in self.exons: for genome_pos in range(estart, eend): if self.strand == '+': self.seqpos2genome[seq_pos] = genome_pos elif self.strand == '-': tmp = self.cds_len - seq_pos - 1 self.seqpos2genome[tmp] = genome_pos seq_pos += 1 # recode 5' splice site locations for i in range(0, self.five_ss_len): seq_pos = self.cds_len + i ss_ix = i // 2 # the ss_ix'th 5'ss starting from upstream tx pos_in_ss = i % 2 # whether first/second nuc in splice site # determine genome coordinates for 5' splice site if self.strand == '+': self.seqpos2genome[seq_pos] = self.exons[ss_ix][1] + pos_in_ss else: exon_pos = -1 - ss_ix self.seqpos2genome[seq_pos] = self.exons[exon_pos][0] - pos_in_ss - 1 # recode 3' splice site locations for i in range(0, self.three_ss_len): seq_pos = self.cds_len + self.five_ss_len + i ss_ix = i // 2 # the ss_ix'th 3'ss starting from upstream tx pos_in_ss = i % 2 # whether first/second nuc in splice site # determine genome coordinates for 3' splice site if self.strand == '+': self.seqpos2genome[seq_pos] = self.exons[ss_ix+1][0] - 2 + pos_in_ss else: exon_pos = -1 - ss_ix self.seqpos2genome[seq_pos] = self.exons[exon_pos-1][1] + 1 - pos_in_ss
Creates the self.seqpos2genome dictionary that converts positions relative to the sequence to genome coordinates.
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def query_position(self, strand, chr, genome_coord): """Provides the relative position on the coding sequence for a given genomic position. Parameters ---------- chr : str chromosome, provided to check validity of query genome_coord : int 0-based position for mutation, actually used to get relative coding pos Returns ------- pos : int or None position of mutation in coding sequence, returns None if mutation does not match region found in self.exons """ # first check if valid pos = None # initialize to invalid pos if chr != self.chrom: #logger.debug('Wrong chromosome queried. You provided {0} but gene is ' #'on {1}.'.format(chr, self.chrom)) # return pos pass if type(genome_coord) is list: # handle case for indels pos_left = self.query_position(strand, chr, genome_coord[0]) pos_right = self.query_position(strand, chr, genome_coord[1]) if pos_left is not None or pos_right is not None: return [pos_left, pos_right] else: return None # return position if contained within coding region or splice site for i, (estart, eend) in enumerate(self.exons): # in coding region if estart <= genome_coord < eend: if strand == '+': prev_lens = sum(self.exon_lens[:i]) # previous exon lengths pos = prev_lens + (genome_coord - estart) elif strand == '-': prev_lens = sum(self.exon_lens[:i]) # previous exon lengths pos = prev_lens + (genome_coord - estart) pos = self.cds_len - pos - 1 # flip coords because neg strand return pos # in splice site elif (eend <= genome_coord < eend + 2) and i != self.num_exons-1: if strand == '+': pos = self.cds_len + 2*i + (genome_coord - eend) elif strand == '-': pos = self.cds_len + self.five_ss_len + 2*(self.num_exons-(i+2)) + (genome_coord - eend) return pos # in splice site elif (estart - 2 <= genome_coord < estart) and i != 0: if strand == '-': pos = self.cds_len + 2*(self.num_exons-(i+2)) + (genome_coord - (estart - 2)) elif strand == '+': pos = self.cds_len + self.five_ss_len + 2*(i-1) + (genome_coord - (estart - 2)) return pos return pos
Provides the relative position on the coding sequence for a given genomic position. Parameters ---------- chr : str chromosome, provided to check validity of query genome_coord : int 0-based position for mutation, actually used to get relative coding pos Returns ------- pos : int or None position of mutation in coding sequence, returns None if mutation does not match region found in self.exons
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def start_logging(log_file='', log_level='INFO', verbose=False): """Start logging information into the log directory. If os.devnull is specified as the log_file then the log file will not actually be written to a file. """ if not log_file: # create log directory if it doesn't exist log_dir = os.path.abspath('log') + '/' if not os.path.isdir(log_dir): os.mkdir(log_dir) # path to new log file log_file = log_dir + 'log.run.' + str(datetime.datetime.now()).replace(':', '.') + '.txt' # logger options lvl = logging.DEBUG if log_level.upper() == 'DEBUG' else logging.INFO # ignore warnings if not in debug if log_level.upper() != 'DEBUG': warnings.filterwarnings('ignore') # define logging format if verbose: myformat = '%(asctime)s - %(name)s - %(levelname)s \n>>> %(message)s' else: myformat = '%(message)s' # create logger if not log_file == 'stdout': # normal logging to a regular file logging.basicConfig(level=lvl, format=myformat, filename=log_file, filemode='w') else: # logging to stdout root = logging.getLogger() root.setLevel(lvl) stdout_stream = logging.StreamHandler(sys.stdout) stdout_stream.setLevel(lvl) formatter = logging.Formatter(myformat) stdout_stream.setFormatter(formatter) root.addHandler(stdout_stream) root.propagate = True
Start logging information into the log directory. If os.devnull is specified as the log_file then the log file will not actually be written to a file.
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def log_error_decorator(f): """Writes exception to log file if occured in decorated function. This decorator wrapper is needed for multiprocess logging since otherwise the python multiprocessing module will obscure the actual line of the error. """ @wraps(f) def wrapper(*args, **kwds): try: result = f(*args, **kwds) return result except KeyboardInterrupt: logger.info('Ctrl-C stopped a process.') except Exception as e: logger.exception(e) raise return wrapper
Writes exception to log file if occured in decorated function. This decorator wrapper is needed for multiprocess logging since otherwise the python multiprocessing module will obscure the actual line of the error.
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def filter_list(mylist, bad_ixs): """Removes indices from a list. All elements in bad_ixs will be removed from the list. Parameters ---------- mylist : list list to filter out specific indices bad_ixs : list of ints indices to remove from list Returns ------- mylist : list list with elements filtered out """ # indices need to be in reverse order for filtering # to prevent .pop() from yielding eroneous results bad_ixs = sorted(bad_ixs, reverse=True) for i in bad_ixs: mylist.pop(i) return mylist
Removes indices from a list. All elements in bad_ixs will be removed from the list. Parameters ---------- mylist : list list to filter out specific indices bad_ixs : list of ints indices to remove from list Returns ------- mylist : list list with elements filtered out
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def rev_comp(seq): """Get reverse complement of sequence. rev_comp will maintain the case of the sequence. Parameters ---------- seq : str nucleotide sequence. valid {a, c, t, g, n} Returns ------- rev_comp_seq : str reverse complement of sequence """ rev_seq = seq[::-1] rev_comp_seq = ''.join([base_pairing[s] for s in rev_seq]) return rev_comp_seq
Get reverse complement of sequence. rev_comp will maintain the case of the sequence. Parameters ---------- seq : str nucleotide sequence. valid {a, c, t, g, n} Returns ------- rev_comp_seq : str reverse complement of sequence
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def bed_generator(bed_path): """Iterates through a BED file yielding parsed BED lines. Parameters ---------- bed_path : str path to BED file Yields ------ BedLine(line) : BedLine A BedLine object which has parsed the individual line in a BED file. """ with open(bed_path) as handle: bed_reader = csv.reader(handle, delimiter='\t') for line in bed_reader: yield BedLine(line)
Iterates through a BED file yielding parsed BED lines. Parameters ---------- bed_path : str path to BED file Yields ------ BedLine(line) : BedLine A BedLine object which has parsed the individual line in a BED file.
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def read_bed(file_path, restricted_genes=None): """Reads BED file and populates a dictionary separating genes by chromosome. Parameters ---------- file_path : str path to BED file filtered_genes: list list of gene names to not use Returns ------- bed_dict: dict dictionary mapping chromosome keys to a list of BED lines """ # read in entire bed file into a dict with keys as chromsomes bed_dict = OrderedDict() for bed_row in bed_generator(file_path): is_restrict_flag = restricted_genes is None or bed_row.gene_name in restricted_genes if is_restrict_flag: bed_dict.setdefault(bed_row.chrom, []) bed_dict[bed_row.chrom].append(bed_row) sort_chroms = sorted(bed_dict.keys(), key=lambda x: len(bed_dict[x]), reverse=True) bed_dict = OrderedDict((chrom, bed_dict[chrom]) for chrom in sort_chroms) return bed_dict
Reads BED file and populates a dictionary separating genes by chromosome. Parameters ---------- file_path : str path to BED file filtered_genes: list list of gene names to not use Returns ------- bed_dict: dict dictionary mapping chromosome keys to a list of BED lines
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def _fix_mutation_df(mutation_df, only_unique=False): """Drops invalid mutations and corrects for 1-based coordinates. TODO: Be smarter about what coordinate system is put in the provided mutations. Parameters ---------- mutation_df : pd.DataFrame user provided mutations only_unique : bool flag indicating whether only unique mutations for each tumor sample should be kept. This avoids issues when the same mutation has duplicate reportings. Returns ------- mutation_df : pd.DataFrame mutations filtered for being valid and correct mutation type. Also converted 1-base coordinates to 0-based. """ # only keep allowed mutation types orig_len = len(mutation_df) # number of mutations before filtering mutation_df = mutation_df[mutation_df.Variant_Classification.isin(variant_snv)] # only keep SNV type_len = len(mutation_df) # number of mutations after filtering based on mut type # log the number of dropped mutations log_msg = ('Dropped {num_dropped} mutations after only keeping ' '{mut_types}. Indels are processed separately.'.format(num_dropped=orig_len-type_len, mut_types=', '.join(variant_snv))) logger.info(log_msg) # check if mutations are valid SNVs valid_nuc_flag = (mutation_df['Reference_Allele'].apply(is_valid_nuc) & \ mutation_df['Tumor_Allele'].apply(is_valid_nuc)) mutation_df = mutation_df[valid_nuc_flag] # filter bad lines mutation_df = mutation_df[mutation_df['Tumor_Allele'].apply(lambda x: len(x)==1)] mutation_df = mutation_df[mutation_df['Reference_Allele'].apply(lambda x: len(x)==1)] valid_len = len(mutation_df) # log the number of dropped mutations log_msg = ('Dropped {num_dropped} mutations after only keeping ' 'valid SNVs'.format(num_dropped=type_len-valid_len)) logger.info(log_msg) # drop duplicate mutations if only_unique: dup_cols = ['Tumor_Sample', 'Chromosome', 'Start_Position', 'End_Position', 'Reference_Allele', 'Tumor_Allele'] mutation_df = mutation_df.drop_duplicates(subset=dup_cols) # log results of de-duplication dedup_len = len(mutation_df) log_msg = ('Dropped {num_dropped} mutations when removing ' 'duplicates'.format(num_dropped=valid_len-dedup_len)) logger.info(log_msg) # add dummy Protein_Change or Tumor_Type columns if not provided # in file if 'Tumor_Type' not in mutation_df.columns: mutation_df['Tumor_Type'] = '' if 'Protein_Change' not in mutation_df.columns: mutation_df['Protein_Change'] = '' # correct for 1-based coordinates mutation_df['Start_Position'] = mutation_df['Start_Position'].astype(int) - 1 return mutation_df
Drops invalid mutations and corrects for 1-based coordinates. TODO: Be smarter about what coordinate system is put in the provided mutations. Parameters ---------- mutation_df : pd.DataFrame user provided mutations only_unique : bool flag indicating whether only unique mutations for each tumor sample should be kept. This avoids issues when the same mutation has duplicate reportings. Returns ------- mutation_df : pd.DataFrame mutations filtered for being valid and correct mutation type. Also converted 1-base coordinates to 0-based.
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def calc_windowed_sum(aa_mut_pos, germ_aa, somatic_aa, window=[3]): """Calculate the sum of mutations within a window around a particular mutated codon. Parameters ---------- aa_mut_pos : list list of mutated amino acid positions germ_aa : list Reference amino acid somatic_aa : list Somatic amino acid (if missense) window : list List of windows to calculate for Returns ------- pos_ctr : dict dictionary of mutated positions (key) with associated counts (value) pos_sum : dict of dict Window size as first key points to dictionary of mutated positions (key) with associated mutation count within the window size (value) """ pos_ctr, pos_sum = {}, {w: {} for w in window} num_pos = len(aa_mut_pos) # figure out the missense mutations for i in range(num_pos): pos = aa_mut_pos[i] # make sure mutation is missense if germ_aa[i] and somatic_aa[i] and germ_aa[i] != '*' and \ somatic_aa[i] != '*' and germ_aa[i] != somatic_aa[i]: # should have a position, but if not skip it if pos is not None: pos_ctr.setdefault(pos, 0) pos_ctr[pos] += 1 # calculate windowed sum pos_list = sorted(pos_ctr.keys()) max_window = max(window) for ix, pos in enumerate(pos_list): tmp_sum = {w: 0 for w in window} # go through the same and lower positions for k in reversed(range(ix+1)): pos2 = pos_list[k] if pos2 < pos-max_window: break for w in window: if pos-w <= pos2: tmp_sum[w] += pos_ctr[pos2] # go though the higher positions for l in range(ix+1, len(pos_list)): pos2 = pos_list[l] if pos2 > pos+max_window: break for w in window: if pos2 <= pos+w: tmp_sum[w] += pos_ctr[pos2] # iterate through all other positions #for pos2 in pos_list: #for w in window: #if pos-w <= pos2 <= pos+w: #tmp_sum[w] += pos_ctr[pos2] # update windowed counts for w in window: pos_sum[w][pos] = tmp_sum[w] return pos_ctr, pos_sum
Calculate the sum of mutations within a window around a particular mutated codon. Parameters ---------- aa_mut_pos : list list of mutated amino acid positions germ_aa : list Reference amino acid somatic_aa : list Somatic amino acid (if missense) window : list List of windows to calculate for Returns ------- pos_ctr : dict dictionary of mutated positions (key) with associated counts (value) pos_sum : dict of dict Window size as first key points to dictionary of mutated positions (key) with associated mutation count within the window size (value)
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def get_all_context_names(context_num): """Based on the nucleotide base context number, return a list of strings representing each context. Parameters ---------- context_num : int number representing the amount of nucleotide base context to use. Returns ------- a list of strings containing the names of the base contexts """ if context_num == 0: return ['None'] elif context_num == 1: return ['A', 'C', 'T', 'G'] elif context_num == 1.5: return ['C*pG', 'CpG*', 'TpC*', 'G*pA', 'A', 'C', 'T', 'G'] elif context_num == 2: dinucs = list(set( [d1+d2 for d1 in 'ACTG' for d2 in 'ACTG'] )) return dinucs elif context_num == 3: trinucs = list(set( [t1+t2+t3 for t1 in 'ACTG' for t2 in 'ACTG' for t3 in 'ACTG'] )) return trinucs
Based on the nucleotide base context number, return a list of strings representing each context. Parameters ---------- context_num : int number representing the amount of nucleotide base context to use. Returns ------- a list of strings containing the names of the base contexts
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def get_chasm_context(tri_nuc): """Returns the mutation context acording to CHASM. For more information about CHASM's mutation context, look at http://wiki.chasmsoftware.org/index.php/CHASM_Overview. Essentially CHASM uses a few specified di-nucleotide contexts followed by single nucleotide context. Parameters ---------- tri_nuc : str three nucleotide string with mutated base in the middle. Returns ------- chasm context : str a string representing the context used in CHASM """ # check if string is correct length if len(tri_nuc) != 3: raise ValueError('Chasm context requires a three nucleotide string ' '(Provided: "{0}")'.format(tri_nuc)) # try dinuc context if found if tri_nuc[1:] == 'CG': return 'C*pG' elif tri_nuc[:2] == 'CG': return 'CpG*' elif tri_nuc[:2] == 'TC': return 'TpC*' elif tri_nuc[1:] == 'GA': return 'G*pA' else: # just return single nuc context return tri_nuc[1]
Returns the mutation context acording to CHASM. For more information about CHASM's mutation context, look at http://wiki.chasmsoftware.org/index.php/CHASM_Overview. Essentially CHASM uses a few specified di-nucleotide contexts followed by single nucleotide context. Parameters ---------- tri_nuc : str three nucleotide string with mutated base in the middle. Returns ------- chasm context : str a string representing the context used in CHASM
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def get_aa_mut_info(coding_pos, somatic_base, gene_seq): """Retrieves relevant information about the effect of a somatic SNV on the amino acid of a gene. Information includes the germline codon, somatic codon, codon position, germline AA, and somatic AA. Parameters ---------- coding_pos : iterable of ints Contains the base position (0-based) of the mutations somatic_base : list of str Contains the somatic nucleotide for the mutations gene_seq : GeneSequence gene sequence Returns ------- aa_info : dict information about the somatic mutation effect on AA's """ # if no mutations return empty result if not somatic_base: aa_info = {'Reference Codon': [], 'Somatic Codon': [], 'Codon Pos': [], 'Reference Nuc': [], 'Reference AA': [], 'Somatic AA': []} return aa_info # get codon information into three lists ref_codon, codon_pos, pos_in_codon, ref_nuc = zip(*[cutils.pos_to_codon(gene_seq, p) for p in coding_pos]) ref_codon, codon_pos, pos_in_codon, ref_nuc = list(ref_codon), list(codon_pos), list(pos_in_codon), list(ref_nuc) # construct codons for mutations mut_codon = [(list(x) if x != 'Splice_Site' else []) for x in ref_codon] for i in range(len(mut_codon)): # splice site mutations are not in a codon, so skip such mutations to # prevent an error if pos_in_codon[i] is not None: pc = pos_in_codon[i] mut_codon[i][pc] = somatic_base[i] mut_codon = [(''.join(x) if x else 'Splice_Site') for x in mut_codon] # output resulting info aa_info = {'Reference Codon': ref_codon, 'Somatic Codon': mut_codon, 'Codon Pos': codon_pos, 'Reference Nuc': ref_nuc, 'Reference AA': [(utils.codon_table[r] if (r in utils.codon_table) else None) for r in ref_codon], 'Somatic AA': [(utils.codon_table[s] if (s in utils.codon_table) else None) for s in mut_codon]} return aa_info
Retrieves relevant information about the effect of a somatic SNV on the amino acid of a gene. Information includes the germline codon, somatic codon, codon position, germline AA, and somatic AA. Parameters ---------- coding_pos : iterable of ints Contains the base position (0-based) of the mutations somatic_base : list of str Contains the somatic nucleotide for the mutations gene_seq : GeneSequence gene sequence Returns ------- aa_info : dict information about the somatic mutation effect on AA's
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def handle_tsg_results(permutation_result): """Handles result from TSG results. Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save """ permutation_df = pd.DataFrame(sorted(permutation_result, key=lambda x: x[2] if x[2] is not None else 1.1), columns=['gene', 'inactivating count', 'inactivating p-value', 'Total SNV Mutations', 'SNVs Unmapped to Ref Tx']) permutation_df['inactivating p-value'] = permutation_df['inactivating p-value'].astype('float') tmp_df = permutation_df[permutation_df['inactivating p-value'].notnull()] # get benjamani hochberg adjusted p-values permutation_df['inactivating BH q-value'] = np.nan permutation_df.loc[tmp_df.index, 'inactivating BH q-value'] = mypval.bh_fdr(tmp_df['inactivating p-value']) # sort output by p-value. due to no option to specify NaN order in # sort, the df needs to sorted descendingly and then flipped permutation_df = permutation_df.sort_values(by='inactivating p-value', ascending=False) permutation_df = permutation_df.reindex(index=permutation_df.index[::-1]) # order result permutation_df = permutation_df.set_index('gene', drop=False) col_order = ['gene', 'Total SNV Mutations', 'SNVs Unmapped to Ref Tx', #'Total Frameshift Mutations', 'Frameshifts Unmapped to Ref Tx', 'inactivating count', 'inactivating p-value', 'inactivating BH q-value'] return permutation_df[col_order]
Handles result from TSG results. Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save
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def handle_oncogene_results(permutation_result, num_permutations): """Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save """ mycols = ['gene', 'num recurrent', 'position entropy', 'mean vest score', 'entropy p-value', 'vest p-value', 'Total Mutations', 'Unmapped to Ref Tx'] permutation_df = pd.DataFrame(permutation_result, columns=mycols) # get benjamani hochberg adjusted p-values permutation_df['entropy BH q-value'] = mypval.bh_fdr(permutation_df['entropy p-value']) permutation_df['vest BH q-value'] = mypval.bh_fdr(permutation_df['vest p-value']) # combine p-values permutation_df['tmp entropy p-value'] = permutation_df['entropy p-value'] permutation_df['tmp vest p-value'] = permutation_df['vest p-value'] permutation_df.loc[permutation_df['entropy p-value']==0, 'tmp entropy p-value'] = 1. / num_permutations permutation_df.loc[permutation_df['vest p-value']==0, 'tmp vest p-value'] = 1. / num_permutations permutation_df['combined p-value'] = permutation_df[['entropy p-value', 'vest p-value']].apply(mypval.fishers_method, axis=1) permutation_df['combined BH q-value'] = mypval.bh_fdr(permutation_df['combined p-value']) del permutation_df['tmp vest p-value'] del permutation_df['tmp entropy p-value'] # order output permutation_df = permutation_df.set_index('gene', drop=False) # make sure genes are indices permutation_df['num recurrent'] = permutation_df['num recurrent'].fillna(-1).astype(int) # fix dtype isssue col_order = ['gene', 'Total Mutations', 'Unmapped to Ref Tx', 'num recurrent', 'position entropy', 'mean vest score', 'entropy p-value', 'vest p-value', 'combined p-value', 'entropy BH q-value', 'vest BH q-value', 'combined BH q-value'] permutation_df = permutation_df.sort_values(by=['combined p-value']) return permutation_df[col_order]
Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save
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def handle_hotmaps_results(permutation_result): """Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save """ if len(permutation_result[0]) == 6: mycols = ['gene', 'window length', 'codon position', 'mutation count', 'windowed sum', 'p-value'] else: mycols = ['gene', 'window length', 'codon position', 'index', 'mutation count', 'windowed sum', 'p-value'] permutation_df = pd.DataFrame(permutation_result, columns=mycols) # get benjamani hochberg adjusted p-values permutation_df['q-value'] = 1 for w in permutation_df['window length'].unique(): is_window = permutation_df['window length'] == w permutation_df.loc[is_window, 'q-value'] = mypval.bh_fdr(permutation_df.loc[is_window, 'p-value']) #permutation_df['q-value'] = mypval.bh_fdr(permutation_df['p-value']) # order output #permutation_df = permutation_df.set_index('gene', drop=False) # make sure genes are indices col_order = mycols + ['q-value'] permutation_df = permutation_df.sort_values(by=['window length', 'p-value']) return permutation_df[col_order]
Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save
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def handle_protein_results(permutation_result): """Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save """ mycols = ['gene', 'num recurrent', 'normalized graph-smoothed position entropy', 'normalized graph-smoothed position entropy p-value', 'Total Mutations', 'Unmapped to Ref Tx'] permutation_df = pd.DataFrame(permutation_result, columns=mycols) # get benjamani hochberg adjusted p-values permutation_df['normalized graph-smoothed position entropy BH q-value'] = mypval.bh_fdr(permutation_df['normalized graph-smoothed position entropy p-value']) # order output permutation_df = permutation_df.set_index('gene', drop=False) # make sure genes are indices col_order = ['gene', 'Total Mutations', 'Unmapped to Ref Tx', 'num recurrent', 'normalized graph-smoothed position entropy', 'normalized graph-smoothed position entropy p-value', 'normalized graph-smoothed position entropy BH q-value'] permutation_df = permutation_df.sort_values(by=['normalized graph-smoothed position entropy p-value']) return permutation_df[col_order]
Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save
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def handle_effect_results(permutation_result): """Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save """ mycols = ['gene', 'num recurrent', 'num inactivating', 'entropy-on-effect', 'entropy-on-effect p-value', 'Total Mutations', 'Unmapped to Ref Tx'] permutation_df = pd.DataFrame(sorted(permutation_result, key=lambda x: x[4] if x[4] is not None else 1.1), columns=mycols) # get benjamani hochberg adjusted p-values permutation_df['entropy-on-effect BH q-value'] = mypval.bh_fdr(permutation_df['entropy-on-effect p-value']) # order output permutation_df = permutation_df.set_index('gene', drop=False) # make sure genes are indices permutation_df['num recurrent'] = permutation_df['num recurrent'].fillna(-1).astype(int) # fix dtype isssue col_order = ['gene', 'Total Mutations', 'Unmapped to Ref Tx', 'num recurrent', 'num inactivating', 'entropy-on-effect', 'entropy-on-effect p-value', 'entropy-on-effect BH q-value'] return permutation_df[col_order]
Takes in output from multiprocess_permutation function and converts to a better formatted dataframe. Parameters ---------- permutation_result : list output from multiprocess_permutation Returns ------- permutation_df : pd.DataFrame formatted output suitable to save
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def get_frameshift_info(fs_df, bins): """Counts frameshifts stratified by a given length. Parameters ---------- fs_df : pd.DataFrame indel mutations from non-coding portion bins : int number of different length categories for frameshifts Returns ------- indel_len : list length of specific frameshift length category num_indels : list number of frameshifts matchin indel_len """ fs_df = compute_indel_length(fs_df) # count the number INDELs with length non-dividable by 3 num_indels = [] indel_len = [] num_categories = 0 i = 1 while(num_categories<bins): # not inframe length indel if i%3: if num_categories != bins-1: tmp_num = len(fs_df[fs_df['indel len']==i]) else: tmp_num = len(fs_df[(fs_df['indel len']>=i) & ((fs_df['indel len']%3)>0)]) num_indels.append(tmp_num) indel_len.append(i) num_categories += 1 i += 1 return indel_len, num_indels
Counts frameshifts stratified by a given length. Parameters ---------- fs_df : pd.DataFrame indel mutations from non-coding portion bins : int number of different length categories for frameshifts Returns ------- indel_len : list length of specific frameshift length category num_indels : list number of frameshifts matchin indel_len
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def set_mutation_type(self, mut_type=''): """Sets the mutation type attribute to a single label based on attribute flags. Kwargs: mut_type (str): value to set self.mut_type """ if mut_type: # user specifies a mutation type self.mutation_type = mut_type else: # mutation type is taken from object attributes if not self.is_valid: # does not correctly fall into a category self.mutation_type = 'not valid' elif self.unknown_effect: self.mutation_type = 'unknown effect' elif self.is_no_protein: self.mutation_type = 'no protein' elif self.is_missing_info: # mutation has a ? self.mutation_type = 'missing' else: # valid mutation type to be counted if self.is_lost_stop: self.mutation_type = 'Nonstop_Mutation' elif self.is_lost_start: self.mutation_type = 'Translation_Start_Site' elif self.is_synonymous: # synonymous must go before missense since mutations # can be categorized as synonymous and missense. Although # in reality such cases are actually synonymous and not # missense mutations. self.mutation_type = 'Silent' elif self.is_missense: self.mutation_type = 'Missense_Mutation' elif self.is_indel: self.mutation_type = 'In_Frame_Indel' elif self.is_nonsense_mutation: self.mutation_type = 'Nonsense_Mutation' elif self.is_frame_shift: self.mutation_type = 'Frame_Shift_Indel'
Sets the mutation type attribute to a single label based on attribute flags. Kwargs: mut_type (str): value to set self.mut_type
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def set_amino_acid(self, aa): """Set amino acid change and position.""" aa = aa.upper() # make sure it is upper case aa = aa[2:] if aa.startswith('P.') else aa # strip "p." self.__set_mutation_status() # set flags detailing the type of mutation self.__parse_hgvs_syntax(aa)
Set amino acid change and position.
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def __set_mutation_type(self, hgvs_string): """Interpret the mutation type (missense, etc.) and set appropriate flags. Args: hgvs_string (str): hgvs syntax with "p." removed """ self.__set_lost_stop_status(hgvs_string) self.__set_lost_start_status(hgvs_string) self.__set_missense_status(hgvs_string) # missense mutations self.__set_indel_status() # indel mutations self.__set_frame_shift_status() # check for fs self.__set_premature_stop_codon_status(hgvs_string)
Interpret the mutation type (missense, etc.) and set appropriate flags. Args: hgvs_string (str): hgvs syntax with "p." removed
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def __set_missense_status(self, hgvs_string): """Sets the self.is_missense flag.""" # set missense status if re.search('^[A-Z?]\d+[A-Z?]$', hgvs_string): self.is_missense = True self.is_non_silent = True else: self.is_missense = False
Sets the self.is_missense flag.
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def __set_lost_start_status(self, hgvs_string): """Sets the self.is_lost_start flag.""" # set is lost start status mymatch = re.search('^([A-Z?])(\d+)([A-Z?])$', hgvs_string) if mymatch: grps = mymatch.groups() if int(grps[1]) == 1 and grps[0] != grps[2]: self.is_lost_start = True self.is_non_silent = True else: self.is_lost_start = False else: self.is_lost_start = False
Sets the self.is_lost_start flag.
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def __set_frame_shift_status(self): """Check for frame shift and set the self.is_frame_shift flag.""" if 'fs' in self.hgvs_original: self.is_frame_shift = True self.is_non_silent = True elif re.search('[A-Z]\d+[A-Z]+\*', self.hgvs_original): # it looks like some mutations dont follow the convention # of using 'fs' to indicate frame shift self.is_frame_shift = True self.is_non_silent = True else: self.is_frame_shift = False
Check for frame shift and set the self.is_frame_shift flag.
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def __set_lost_stop_status(self, hgvs_string): """Check if the stop codon was mutated to something other than a stop codon.""" lost_stop_pattern = '^\*\d+[A-Z?]+\*?$' if re.search(lost_stop_pattern, hgvs_string): self.is_lost_stop = True self.is_non_silent = True else: self.is_lost_stop = False
Check if the stop codon was mutated to something other than a stop codon.
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def __set_premature_stop_codon_status(self, hgvs_string): """Set whether there is a premature stop codon.""" if re.search('.+\*(\d+)?$', hgvs_string): self.is_premature_stop_codon = True self.is_non_silent = True # check if it is also a nonsense mutation if hgvs_string.endswith('*'): self.is_nonsense_mutation = True else: self.is_nonsense_mutation = False else: self.is_premature_stop_codon = False self.is_nonsense_mutation = False
Set whether there is a premature stop codon.
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def __set_indel_status(self): """Sets flags related to the mutation being an indel.""" # set indel status if "ins" in self.hgvs_original: # mutation is insertion self.is_insertion = True self.is_deletion = False self.is_indel = True self.is_non_silent = True elif "del" in self.hgvs_original: # mutation is deletion self.is_deletion = True self.is_insertion = False self.is_indel = True self.is_non_silent = True else: # not an indel self.is_deletion = False self.is_insertion = False self.is_indel = False
Sets flags related to the mutation being an indel.
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def __set_unkown_effect(self, hgvs_string): """Sets a flag for unkown effect according to HGVS syntax. The COSMIC database also uses unconventional questionmarks to denote missing information. Args: hgvs_string (str): hgvs syntax with "p." removed """ # Standard use by HGVS of indicating unknown effect. unknown_effect_list = ['?', '(=)', '='] # unknown effect symbols if hgvs_string in unknown_effect_list: self.unknown_effect = True elif "(" in hgvs_string: # parethesis in HGVS indicate expected outcomes self.unknown_effect = True else: self.unknown_effect = False # detect if there are missing information. commonly COSMIC will # have insertions with p.?_?ins? or deleteions with ?del indicating # missing information. if "?" in hgvs_string: self.is_missing_info = True else: self.is_missing_info = False
Sets a flag for unkown effect according to HGVS syntax. The COSMIC database also uses unconventional questionmarks to denote missing information. Args: hgvs_string (str): hgvs syntax with "p." removed
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def __set_no_protein(self, hgvs_string): """Set a flag for no protein expected. ("p.0" or "p.0?") Args: hgvs_string (str): hgvs syntax with "p." removed """ no_protein_list = ['0', '0?'] # no protein symbols if hgvs_string in no_protein_list: self.is_no_protein = True self.is_non_silent = True else: self.is_no_protein = False
Set a flag for no protein expected. ("p.0" or "p.0?") Args: hgvs_string (str): hgvs syntax with "p." removed
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def __parse_hgvs_syntax(self, aa_hgvs): """Convert HGVS syntax for amino acid change into attributes. Specific details of the mutation are stored in attributes like self.intial (prior to mutation), sel.pos (mutation position), self.mutated (mutation), and self.stop_pos (position of stop codon, if any). Args: aa_hgvs (str): amino acid string following HGVS syntax """ self.is_valid = True # assume initially the syntax is legitimate self.is_synonymous = False # assume not synonymous until proven if self.unknown_effect or self.is_no_protein: # unknown effect from mutation. usually denoted as p.? self.pos = None pass elif self.is_lost_stop: self.initial = aa_hgvs[0] self.mutated = re.findall('([A-Z?*]+)$', aa_hgvs)[0] self.pos = int(re.findall('^\*(\d+)', aa_hgvs)[0]) self.stop_pos = None elif self.is_lost_start: self.initial = aa_hgvs[0] self.mutated = aa_hgvs[-1] self.pos = int(aa_hgvs[1:-1]) elif self.is_missense: self.initial = aa_hgvs[0] self.mutated = aa_hgvs[-1] self.pos = int(aa_hgvs[1:-1]) self.stop_pos = None # not a nonsense mutation if self.initial == self.mutated: self.is_synonymous = True self.is_non_silent = False elif self.mutated == '*': self.is_nonsense_mutation = True elif self.is_indel: if self.is_insertion: if not self.is_missing_info: self.initial = re.findall('([A-Z])\d+', aa_hgvs)[:2] # first two self.pos = tuple(map(int, re.findall('[A-Z](\d+)', aa_hgvs)[:2])) # first two self.mutated = re.findall('(?<=INS)[A-Z0-9?*]+', aa_hgvs)[0] self.mutated = self.mutated.strip('?') # remove the missing info '?' else: self.initial = '' self.pos = tuple() self.mutated = '' elif self.is_deletion: if not self.is_missing_info: self.initial = re.findall('([A-Z])\d+', aa_hgvs) self.pos = tuple(map(int, re.findall('[A-Z](\d+)', aa_hgvs))) self.mutated = re.findall('(?<=DEL)[A-Z]*', aa_hgvs)[0] else: self.initial = '' self.pos = tuple() self.mutated = '' elif self.is_frame_shift: self.initial = aa_hgvs[0] self.mutated = '' try: self.pos = int(re.findall('[A-Z*](\d+)', aa_hgvs)[0]) if self.is_premature_stop_codon: self.stop_pos = int(re.findall('\*>?(\d+)$', aa_hgvs)[0]) else: self.stop_pos = None except IndexError: # unconventional usage of indicating frameshifts will cause # index errors. For example, in some cases 'fs' is not used. # In other cases, either amino acids were not included or # just designated as a '?' self.logger.debug('(Parsing-Problem) frame shift hgvs string: "%s"' % aa_hgvs) self.pos = None self.stop_pos = None self.is_missing_info = True elif self.is_nonsense_mutation: self.initial = aa_hgvs[0] self.mutated = '*' # there is actually a stop codon self.stop_pos = 0 # indicates same position is stop codon try: self.pos = int(aa_hgvs[1:-1]) except ValueError: # wierd error of p.E217>D* self.is_valid = False self.pos = None self.logger.debug('(Parsing-Problem) Invalid HGVS Amino Acid ' 'syntax: ' + aa_hgvs) if self.initial == self.mutated: # classify nonsense-to-nonsense mutations as synonymous self.is_synonymous = True self.is_non_silent = False else: self.is_valid = False # did not match any of the possible cases self.logger.debug('(Parsing-Problem) Invalid HGVS Amino Acid ' 'syntax: ' + aa_hgvs)
Convert HGVS syntax for amino acid change into attributes. Specific details of the mutation are stored in attributes like self.intial (prior to mutation), sel.pos (mutation position), self.mutated (mutation), and self.stop_pos (position of stop codon, if any). Args: aa_hgvs (str): amino acid string following HGVS syntax
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def deleterious_permutation(obs_del, context_counts, context_to_mut, seq_context, gene_seq, num_permutations=10000, stop_criteria=100, pseudo_count=0, max_batch=25000): """Performs null-permutations for deleterious mutation statistics in a single gene. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null pseudo_count : int, default: 0 Pseudo-count for number of deleterious mutations for each permutation of the null distribution. Increasing pseudo_count makes the statistical test more stringent. Returns ------- del_count_list : list list of deleterious mutation counts under the null """ mycontexts = context_counts.index.tolist() somatic_base = [base for one_context in mycontexts for base in context_to_mut[one_context]] # calculate the # of batches for simulations max_batch = min(num_permutations, max_batch) num_batches = num_permutations // max_batch remainder = num_permutations % max_batch batch_sizes = [max_batch] * num_batches if remainder: batch_sizes += [remainder] num_sim = 0 null_del_ct = 0 for j, batch_size in enumerate(batch_sizes): # stop iterations if reached sufficient precision if null_del_ct >= stop_criteria: #j = j - 1 break # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), batch_size) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # determine result of random positions for i, row in enumerate(tmp_mut_pos): # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # calc deleterious mutation info tmp_del_count = cutils.calc_deleterious_info(tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], tmp_mut_info['Codon Pos']) # update empricial null distribution if tmp_del_count >= obs_del: null_del_ct += 1 # stop if reach sufficient precision on p-value if null_del_ct >= stop_criteria: break # update number of simulations num_sim += i + 1 #num_sim = j*max_batch + i+1 del_pval = float(null_del_ct) / (num_sim) return del_pval
Performs null-permutations for deleterious mutation statistics in a single gene. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null pseudo_count : int, default: 0 Pseudo-count for number of deleterious mutations for each permutation of the null distribution. Increasing pseudo_count makes the statistical test more stringent. Returns ------- del_count_list : list list of deleterious mutation counts under the null
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def position_permutation(obs_stat, context_counts, context_to_mut, seq_context, gene_seq, gene_vest=None, num_permutations=10000, stop_criteria=100, pseudo_count=0, max_batch=25000): """Performs null-permutations for position-based mutation statistics in a single gene. Parameters ---------- obs_stat : tuple, (recur ct, entropy, delta entropy, mean vest) tuple containing the observed statistics context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null stop_criteria : int stop after stop_criteria iterations are more significant then the observed statistic. pseudo_count : int, default: 0 Pseudo-count for number of recurrent missense mutations for each permutation for the null distribution. Increasing pseudo_count makes the statistical test more stringent. Returns ------- num_recur_list : list list of recurrent mutation counts under the null entropy_list : list list of position entropy values under the null """ # get contexts and somatic base mycontexts = context_counts.index.tolist() somatic_base = [base for one_context in mycontexts for base in context_to_mut[one_context]] # calculate the # of batches for simulations max_batch = min(num_permutations, max_batch) num_batches = num_permutations // max_batch remainder = num_permutations % max_batch batch_sizes = [max_batch] * num_batches if remainder: batch_sizes += [remainder] obs_recur, obs_ent, obs_delta_ent, obs_vest = obs_stat num_sim = 0 # number of simulations null_num_recur_ct, null_entropy_ct, null_delta_entropy_ct, null_vest_ct = 0, 0, 0, 0 for j, batch_size in enumerate(batch_sizes): # stop iterations if reached sufficient precision if null_vest_ct >= stop_criteria and null_entropy_ct >= stop_criteria: break # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), batch_size) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # calculate position-based statistics as a result of random positions for i, row in enumerate(tmp_mut_pos): # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # calculate position info tmp_recur_ct, tmp_entropy, tmp_delta_entropy, _ = cutils.calc_pos_info(tmp_mut_info['Codon Pos'], tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], pseudo_count=pseudo_count, is_obs=0) # get vest scores if gene_vest: tmp_vest = scores.compute_vest_stat(gene_vest, tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], tmp_mut_info['Codon Pos']) else: tmp_vest = 0.0 # update empirical null distribution counts if tmp_entropy-utils.epsilon <= obs_ent: null_entropy_ct += 1 if tmp_vest+utils.epsilon >= obs_vest: null_vest_ct += 1 # stop iterations if reached sufficient precision if null_vest_ct >= stop_criteria and null_entropy_ct >= stop_criteria: break # update the number of simulations num_sim += i+1 # calculate p-value from empirical null-distribution ent_pval = float(null_entropy_ct) / (num_sim) vest_pval = float(null_vest_ct) / (num_sim) return ent_pval, vest_pval
Performs null-permutations for position-based mutation statistics in a single gene. Parameters ---------- obs_stat : tuple, (recur ct, entropy, delta entropy, mean vest) tuple containing the observed statistics context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null stop_criteria : int stop after stop_criteria iterations are more significant then the observed statistic. pseudo_count : int, default: 0 Pseudo-count for number of recurrent missense mutations for each permutation for the null distribution. Increasing pseudo_count makes the statistical test more stringent. Returns ------- num_recur_list : list list of recurrent mutation counts under the null entropy_list : list list of position entropy values under the null
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def hotmaps_permutation(obs_stat, context_counts, context_to_mut, seq_context, gene_seq, window, num_permutations=10000, stop_criteria=100, max_batch=25000, null_save_path=None): """Performs null-permutations for position-based mutation statistics in a single gene. Parameters ---------- obs_stat : dict dictionary mapping codons to the sum of mutations in a window context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest window : int Number of codons to the left/right of a mutated position to consider in the window num_permutations : int, default: 10000 number of permutations to create for null stop_criteria : int stop after stop_criteria iterations are more significant then the observed statistic. max_batch : int maximum number of whole gene simulations to do at once. For large number of simulations holding a matrix of M x N, where M is the number of mutations and N is the number of simulations, can get quite large. null_save_path : str or None File path to save null distribution. If None, don't save it. Returns ------- pvals : dict Maps mutated codon position to the calculated p-value """ # get contexts and somatic base mycontexts = context_counts.index.tolist() somatic_base = [base for one_context in mycontexts for base in context_to_mut[one_context]] # calculate the # of batches for simulations max_batch = min(num_permutations, max_batch) num_batches = num_permutations // max_batch remainder = num_permutations % max_batch batch_sizes = [max_batch] * num_batches if remainder: batch_sizes += [remainder] # figure out which position has highest value max_key = {w: max(obs_stat[w], key=(lambda key: obs_stat[w][key])) for w in window} # setup null dist counts null_cts = {w: {k: 0 for k in obs_stat[w]} for w in window } # empirical null distribution (saved if file path provided) empirical_null = {w: {} for w in window} num_sim = 0 # number of simulations for j, batch_size in enumerate(batch_sizes): # stop iterations if reached sufficient precision # stop iterations if reached sufficient precision stop_flag = [(null_cts[w][max_key[w]]>=stop_criteria) for w in window] if all(stop_flag): break #if null_cts[max_key] >= stop_criteria: #break # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), batch_size) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # calculate position-based statistics as a result of random positions for i, row in enumerate(tmp_mut_pos): # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # calculate position info tmp_pos, tmp_sim = utils.calc_windowed_sum(tmp_mut_info['Codon Pos'], tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], window) # update the counts when the empirical null passes the observed for tmp_w in tmp_sim: for tmp_key in tmp_sim[tmp_w]: # get mutation count for simulation val = tmp_sim[tmp_w][tmp_key] # add to empirical null distribution empirical_null[tmp_w].setdefault(val, 0) empirical_null[tmp_w][val] += 1 # update counts used for p-value for key in null_cts[tmp_w]: if val >= obs_stat[tmp_w][key]: null_cts[tmp_w][key] += 1 # update the number of simulations num_sim += len(tmp_pos) # stop iterations if reached sufficient precision stop_flag = [(null_cts[w][max_key[w]]>=stop_criteria) for w in window] if all(stop_flag): break # calculate p-value from empirical null-distribution pvals = {w: {k: float(null_cts[w][k]) / (num_sim) for k in obs_stat[w]} for w in window} # save empirical distribution if null_save_path: for w in window: # create null distribution output = [['mutation_count', 'p-value']] sorted_cts = sorted(empirical_null[w].keys()) tmp_sum = 0 for i in range(len(sorted_cts)): tmp_sum += empirical_null[w][sorted_cts[-(i+1)]] tmp_pval = tmp_sum / float(num_sim) output.append([sorted_cts[-(i+1)], tmp_pval]) # save output with open(null_save_path.format(w), 'w') as handle: mywriter = csv.writer(handle, delimiter='\t', lineterminator='\n') mywriter.writerows(output) return pvals
Performs null-permutations for position-based mutation statistics in a single gene. Parameters ---------- obs_stat : dict dictionary mapping codons to the sum of mutations in a window context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest window : int Number of codons to the left/right of a mutated position to consider in the window num_permutations : int, default: 10000 number of permutations to create for null stop_criteria : int stop after stop_criteria iterations are more significant then the observed statistic. max_batch : int maximum number of whole gene simulations to do at once. For large number of simulations holding a matrix of M x N, where M is the number of mutations and N is the number of simulations, can get quite large. null_save_path : str or None File path to save null distribution. If None, don't save it. Returns ------- pvals : dict Maps mutated codon position to the calculated p-value
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def protein_permutation(graph_score, num_codons_obs, context_counts, context_to_mut, seq_context, gene_seq, gene_graph, num_permutations=10000, stop_criteria=100, pseudo_count=0): """Performs null-simulations for position-based mutation statistics in a single gene. Parameters ---------- graph_score : float clustering score for observed data num_codons_obs : int number of codons with missense mutation in observed data context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null stop_criteria : int stop after stop_criteria iterations are more significant then the observed statistic. Returns ------- protein_pval : float p-value for clustering in neighbor graph constructure from protein structures """ # get contexts and somatic base mycontexts = context_counts.index.tolist() somatic_base = [base for one_context in mycontexts for base in context_to_mut[one_context]] # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), num_permutations) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # calculate position-based statistics as a result of random positions null_graph_entropy_ct = 0 coverage_list = [] num_mut_list = [] graph_entropy_list = [] for i, row in enumerate(tmp_mut_pos): # calculate the expected value of the relative increase in coverage if i == stop_criteria-1: rel_inc = [coverage_list[k] / float(num_mut_list[k]) for k in range(stop_criteria-1) if coverage_list[k]] exp_rel_inc = np.mean(rel_inc) # calculate observed statistic if num_codons_obs: obs_stat = graph_score / np.log2(exp_rel_inc*num_codons_obs) else: obs_stat = 1.0 # calculate statistics for simulated data sim_stat_list = [ent / np.log2(exp_rel_inc*num_mut_list[l]) for l, ent in enumerate(graph_entropy_list)] null_graph_entropy_ct = len([s for s in sim_stat_list if s-utils.epsilon <= obs_stat]) # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # calculate position info tmp_tuple = cutils.calc_pos_info(tmp_mut_info['Codon Pos'], tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], pseudo_count=pseudo_count, is_obs=0) _, _, _, tmp_pos_ct = tmp_tuple # record num of mut codons if i < stop_criteria-1: tmp_num_mut_codons = len(tmp_pos_ct) num_mut_list.append(tmp_num_mut_codons) # get entropy on graph-smoothed probability distribution tmp_graph_entropy, tmp_coverage = scores.compute_ng_stat(gene_graph, tmp_pos_ct) # record the "coverage" in the graph if i < stop_criteria-1: coverage_list.append(tmp_coverage) graph_entropy_list.append(tmp_graph_entropy) # update empirical null distribution counts if i >= stop_criteria: #if tmp_graph_entropy-utils.epsilon <= graph_score: if tmp_num_mut_codons: sim_stat = tmp_graph_entropy / np.log2(exp_rel_inc*tmp_num_mut_codons) else: sim_stat = 1.0 # add count if sim_stat-utils.epsilon <= obs_stat: null_graph_entropy_ct += 1 # stop iterations if reached sufficient precision if null_graph_entropy_ct >= stop_criteria: break # calculate p-value from empirical null-distribution protein_pval = float(null_graph_entropy_ct) / (i+1) return protein_pval, obs_stat
Performs null-simulations for position-based mutation statistics in a single gene. Parameters ---------- graph_score : float clustering score for observed data num_codons_obs : int number of codons with missense mutation in observed data context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null stop_criteria : int stop after stop_criteria iterations are more significant then the observed statistic. Returns ------- protein_pval : float p-value for clustering in neighbor graph constructure from protein structures
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def effect_permutation(context_counts, context_to_mut, seq_context, gene_seq, num_permutations=10000, pseudo_count=0): """Performs null-permutations for effect-based mutation statistics in a single gene. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null pseudo_count : int, default: 0 Pseudo-count for number of recurrent missense mutations for each permutation for the null distribution. Increasing pseudo_count makes the statistical test more stringent. Returns ------- effect_entropy_list : list list of entropy of effect values under the null recur_list : list number of recurrent missense mutations inactivating_list : list number of inactivating mutations """ mycontexts = context_counts.index.tolist() somatic_base = [base for one_context in mycontexts for base in context_to_mut[one_context]] # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), num_permutations) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # calculate position-based statistics as a result of random positions effect_entropy_list, recur_list, inactivating_list = [], [], [] for row in tmp_mut_pos: # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # calculate position info tmp_entropy, tmp_recur, tmp_inactivating = cutils.calc_effect_info(tmp_mut_info['Codon Pos'], tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], pseudo_count=pseudo_count, is_obs=0) effect_entropy_list.append(tmp_entropy) recur_list.append(tmp_recur) inactivating_list.append(tmp_inactivating) return effect_entropy_list, recur_list, inactivating_list
Performs null-permutations for effect-based mutation statistics in a single gene. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null pseudo_count : int, default: 0 Pseudo-count for number of recurrent missense mutations for each permutation for the null distribution. Increasing pseudo_count makes the statistical test more stringent. Returns ------- effect_entropy_list : list list of entropy of effect values under the null recur_list : list number of recurrent missense mutations inactivating_list : list number of inactivating mutations
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def non_silent_ratio_permutation(context_counts, context_to_mut, seq_context, gene_seq, num_permutations=10000): """Performs null-permutations for non-silent ratio across all genes. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null Returns ------- non_silent_count_list : list of tuples list of non-silent and silent mutation counts under the null """ mycontexts = context_counts.index.tolist() somatic_base = [base for one_context in mycontexts for base in context_to_mut[one_context]] # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), num_permutations) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # determine result of random positions non_silent_count_list = [] for row in tmp_mut_pos: # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # calc deleterious mutation info tmp_non_silent = cutils.calc_non_silent_info(tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], tmp_mut_info['Codon Pos']) non_silent_count_list.append(tmp_non_silent) return non_silent_count_list
Performs null-permutations for non-silent ratio across all genes. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null Returns ------- non_silent_count_list : list of tuples list of non-silent and silent mutation counts under the null
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def summary_permutation(context_counts, context_to_mut, seq_context, gene_seq, score_dir, num_permutations=10000, min_frac=0.0, min_recur=2, drop_silent=False): """Performs null-permutations and summarizes the results as features over the gene. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null drop_silent : bool, default=False Flage on whether to drop all silent mutations. Some data sources do not report silent mutations, and the simulations should match this. Returns ------- summary_info_list : list of lists list of non-silent and silent mutation counts under the null along with information on recurrent missense counts and missense positional entropy. """ mycontexts = context_counts.index.tolist() somatic_base = [base for one_context in mycontexts for base in context_to_mut[one_context]] # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), num_permutations) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # determine result of random positions gene_name = gene_seq.bed.gene_name gene_len = gene_seq.bed.cds_len summary_info_list = [] for i, row in enumerate(tmp_mut_pos): # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # Get all metrics summarizing each gene tmp_summary = cutils.calc_summary_info(tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], tmp_mut_info['Codon Pos'], gene_name, score_dir, min_frac=min_frac, min_recur=min_recur) # drop silent if needed if drop_silent: # silent mutation count is index 1 tmp_summary[1] = 0 # limit the precision of floats #pos_ent = tmp_summary[-1] #tmp_summary[-1] = '{0:.5f}'.format(pos_ent) summary_info_list.append([gene_name, i+1, gene_len]+tmp_summary) return summary_info_list
Performs null-permutations and summarizes the results as features over the gene. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null drop_silent : bool, default=False Flage on whether to drop all silent mutations. Some data sources do not report silent mutations, and the simulations should match this. Returns ------- summary_info_list : list of lists list of non-silent and silent mutation counts under the null along with information on recurrent missense counts and missense positional entropy.
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def maf_permutation(context_counts, context_to_mut, seq_context, gene_seq, num_permutations=10000, drop_silent=False): """Performs null-permutations across all genes and records the results in a format like a MAF file. This could be useful for examining the null permutations because the alternative approaches always summarize the results. With the simulated null-permutations, novel metrics can be applied to create an empirical null-distribution. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null drop_silent : bool, default=False Flage on whether to drop all silent mutations. Some data sources do not report silent mutations, and the simulations should match this. Returns ------- maf_list : list of tuples list of null mutations with mutation info in a MAF like format """ mycontexts = context_counts.index.tolist() somatic_base, base_context = zip(*[(base, one_context) for one_context in mycontexts for base in context_to_mut[one_context]]) # get random positions determined by sequence context tmp_contxt_pos = seq_context.random_pos(context_counts.iteritems(), num_permutations) tmp_mut_pos = np.hstack(pos_array for base, pos_array in tmp_contxt_pos) # info about gene gene_name = gene_seq.bed.gene_name strand = gene_seq.bed.strand chrom = gene_seq.bed.chrom gene_seq.bed.init_genome_coordinates() # map seq pos to genome # determine result of random positions maf_list = [] for row in tmp_mut_pos: # get genome coordinate pos2genome = np.vectorize(lambda x: gene_seq.bed.seqpos2genome[x]+1) genome_coord = pos2genome(row) # get info about mutations tmp_mut_info = mc.get_aa_mut_info(row, somatic_base, gene_seq) # get string describing variant var_class = cutils.get_variant_classification(tmp_mut_info['Reference AA'], tmp_mut_info['Somatic AA'], tmp_mut_info['Codon Pos']) # prepare output for k, mysomatic_base in enumerate(somatic_base): # format DNA change ref_nuc = tmp_mut_info['Reference Nuc'][k] nuc_pos = row[k] dna_change = 'c.{0}{1}>{2}'.format(ref_nuc, nuc_pos, mysomatic_base) # format protein change ref_aa = tmp_mut_info['Reference AA'][k] somatic_aa = tmp_mut_info['Somatic AA'][k] codon_pos = tmp_mut_info['Codon Pos'][k] protein_change = 'p.{0}{1}{2}'.format(ref_aa, codon_pos, somatic_aa) # reverse complement if on negative strand if strand == '-': ref_nuc = utils.rev_comp(ref_nuc) mysomatic_base = utils.rev_comp(mysomatic_base) # append results if drop_silent and var_class[k].decode() == 'Silent': continue maf_line = [gene_name, strand, chrom, genome_coord[k], genome_coord[k], ref_nuc, mysomatic_base, base_context[k], dna_change, protein_change, var_class[k].decode()] maf_list.append(maf_line) return maf_list
Performs null-permutations across all genes and records the results in a format like a MAF file. This could be useful for examining the null permutations because the alternative approaches always summarize the results. With the simulated null-permutations, novel metrics can be applied to create an empirical null-distribution. Parameters ---------- context_counts : pd.Series number of mutations for each context context_to_mut : dict dictionary mapping nucleotide context to a list of observed somatic base changes. seq_context : SequenceContext Sequence context for the entire gene sequence (regardless of where mutations occur). The nucleotide contexts are identified at positions along the gene. gene_seq : GeneSequence Sequence of gene of interest num_permutations : int, default: 10000 number of permutations to create for null drop_silent : bool, default=False Flage on whether to drop all silent mutations. Some data sources do not report silent mutations, and the simulations should match this. Returns ------- maf_list : list of tuples list of null mutations with mutation info in a MAF like format
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def markdown(value, extensions=settings.MARKDOWN_EXTENSIONS, extension_configs=settings.MARKDOWN_EXTENSION_CONFIGS, safe=False): """ Render markdown over a given value, optionally using varios extensions. Default extensions could be defined which MARKDOWN_EXTENSIONS option. :returns: A rendered markdown """ return mark_safe(markdown_module.markdown( force_text(value), extensions=extensions, extension_configs=extension_configs, safe_mode=safe))
Render markdown over a given value, optionally using varios extensions. Default extensions could be defined which MARKDOWN_EXTENSIONS option. :returns: A rendered markdown
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def editor_js_initialization(selector, **extra_settings): """ Return script tag with initialization code. """ init_template = loader.get_template( settings.MARKDOWN_EDITOR_INIT_TEMPLATE) options = dict( previewParserPath=reverse('django_markdown_preview'), **settings.MARKDOWN_EDITOR_SETTINGS) options.update(extra_settings) ctx = dict( selector=selector, extra_settings=simplejson.dumps(options) ) return init_template.render(ctx)
Return script tag with initialization code.
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def preview(request): """ Render preview page. :returns: A rendered preview """ if settings.MARKDOWN_PROTECT_PREVIEW: user = getattr(request, 'user', None) if not user or not user.is_staff: from django.contrib.auth.views import redirect_to_login return redirect_to_login(request.get_full_path()) return render( request, settings.MARKDOWN_PREVIEW_TEMPLATE, dict( content=request.POST.get('data', 'No content posted'), css=settings.MARKDOWN_STYLE ))
Render preview page. :returns: A rendered preview
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def register(): """ Register markdown for flatpages. """ admin.site.unregister(FlatPage) admin.site.register(FlatPage, LocalFlatPageAdmin)
Register markdown for flatpages.
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def main(argv=None): """Command line options.""" program_name = __programm_name__ program_version = "v%s" % __version__ program_descrption = __programm_description__ try: # Setup argument parser parser = ArgumentParser(prog=program_name, description=program_descrption) parser.add_argument("-v", "--verbose", dest="verbose", action="count", help="run in verbose mode (-vvv for more, -vvvv to enable connection debugging)") parser.add_argument("-s", "--sudo", action="store_true", help="run supervisorctl actions with sudo (nopasswd))") parser.add_argument("-V", "--version", action="version", version=program_version) parser.add_argument("host-pattern", help="A host-pattern usually refers to a group of hosts. For more details, see Ansible documentation about Patterns.") #parser.add_argument("supervisorctl-action", help="A supervisorctl action (and optional argument). For more details, see Supervisor documentation about the available supervisorctl actions.") subparsers = parser.add_subparsers(help="One of the available supervisorctl actions.", dest="supervisorctl-action") subparsers.add_parser("status", help="Get status info of all processes.") subparsers.add_parser("reread", help="Reread the configuration files of supervisord") subparsers.add_parser("reload", help="Restart remote supervisord") subparsers.add_parser("update", help="Reload the configuration files of supervisord and add/remove processes as necessary") start_subparser = subparsers.add_parser("start", help="Start a process by name") start_subparser.add_argument("process-name", help="Name of the process") stop_subparser = subparsers.add_parser("stop", help="Stop a process by name") stop_subparser.add_argument("process-name", help="Name of the process") restart_subparser = subparsers.add_parser("restart", help="Restart a process by name") restart_subparser.add_argument("process-name", help="Name of the process") remove_subparser = subparsers.add_parser("remove", help="Remove a process by name") remove_subparser.add_argument("process-name", help="Name of the process") # Process arguments args = parser.parse_args(argv) verbose = args.verbose host_pattern = getattr(args, "host-pattern") supervisorctl_action = getattr(args, "supervisorctl-action") sudo = args.sudo ansible_executable = "ansible" supervisorctl_executable = "supervisorctl" ansible_action_option = "-a" if sudo: supervisorctl_command = "sudo " + supervisorctl_executable + " " + supervisorctl_action else: supervisorctl_command = supervisorctl_executable + " " + supervisorctl_action if supervisorctl_action not in ['status', 'reread', 'update', 'reload']: supervisorctl_argument = getattr(args, "process-name") supervisorctl_command = supervisorctl_command + ' ' + supervisorctl_argument if verbose >0: verbose_level = "-"+ "v"*verbose print("Verbose mode on: " + verbose_level) print "Parsed arguments:" print args retcode = call([ansible_executable, host_pattern, verbose_level, ansible_action_option, supervisorctl_command,]) else: retcode = call([ansible_executable, host_pattern, ansible_action_option, supervisorctl_command]) return retcode except KeyboardInterrupt: ### handle keyboard interrupt ### return 0
Command line options.
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def markdown(value, arg=None): """ Render markdown over a given value, optionally using varios extensions. Default extensions could be defined which MARKDOWN_EXTENSIONS option. Syntax: :: {{value|markdown}} {{value|markdown:"tables,codehilite"}} :returns: A rendered markdown """ extensions = (arg and arg.split(',')) or settings.MARKDOWN_EXTENSIONS return _markdown(value, extensions=extensions, safe=False)
Render markdown over a given value, optionally using varios extensions. Default extensions could be defined which MARKDOWN_EXTENSIONS option. Syntax: :: {{value|markdown}} {{value|markdown:"tables,codehilite"}} :returns: A rendered markdown
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